Campanula rotundifolia complex

Well, I don’t like it but I should write something about this Campanula, which is offered in the Shop (seeds).

Once upon a time, when life was easier, Campanula rotundifolia L. was considered a single species with a widespread distribution and with many variable forms depending on the geographic range.

The situation is reflected by the many synonyms accumulated over the years for this species including: Campanula alaskana, C. arctica, C. gieseckeana, C. intercedens, C. petiolata and so on….

Time has passed, and like with many other things in life, people from various parts of the world started to look more closely at the harebell, with newer tools and fresh eyes. It is an extremely variable pretty plant, so we cannot blame them.

Among other articles, I also managed to find on-line, a partial treatment for Campanulaceae for Flora of North America, which I recommend to be read entirely.
https://inaturalist.ca/posts/86696-campanula-partial-treatment-for-flora-of-north-america

For those who want a very short resume:
Firstly, it seems that the species has colonized NA in a single event from European ancestors, then it diverged into separate morphotypes.

Campanula rotundifolia populations in North America are represented by diploids, tetraploids and hexaploids which occur in a ‘matrix’ according with S. G. Shetler who studied C. rotundifolia in North America and is of the opinion that “the tetraploids form a ‘groundmass,’ and the others might turn up anywhere”. (!)

He has been able to differentiate four ‘races’ in NA based on plants morphology: the “Alaskan Race”, the “Arctic Race”, the “Eastern Race” and the “Cordilleran Race” (please follow the above link for more explanations and relative descriptions).
Very important I think is this note: “in any particular location plants may be found with any combination of morphologies mentioned above, or populations with the features of one or more “races” may occur together or in the same general area”.

If all these ‘races’ will be formally recognized as distinct species, I don’t know. I notice that on WFO C. rotundifolia and C. gieseckeana are listed as accepted.

I will personally wait for a general consensus and a complete treatment of Campanulaceae published in Flora of North America, so we can follow the traditional keys for identification. We will follow closely this ‘story’ of and wait for the ‘happy ending’.

With this in mind, it is important to note the seeds collection locations.

Regarding the seeds offered in BotanyCa Shop – all seeds offered as Campanula rotundifolia, of any year, were collected in Ontario on the shorelines of Lake Huron (Bruce peninsula and Killarney).

I also grew plants from the latest collection and after a very close look with the measuring tape in hand out in the garden, our current plants/seeds qualify ‘somewhat’ into to the “Eastern race” as described by S. G. Shetler, with the decumbent stems and very particular ‘ribbonlike leaves’  (currently presented as Campanula intercedens on iNaturalist).
https://www.inaturalist.org/observations?place_id=any&taxon_id=905640

I would also note that height and vigor will vary greatly depending on the garden location: shorter, more upright plants in a full sun/dry location versus a much taller plant in part-shade and more moisture.

So, for now I changed the title in the Shop to:
Campanula rotundifolia L.– Killarney, ON
somewhat equivalent to Campanula intercedens Witasek
If you grew plants from seeds purchased in the last two years from BotanyCa Shop, please mark this change in your labels/records.