Friday’s seeds

It is time to get back to the routine :) We’ll start slowly with a known species – Glaucidium palmatum, Fam. Ranunculaceae (well, there were debates, but it is still there).

The first fruits are ready, more to come, so just a short stop to admire how neatly the seeds are arranged in the follicle. Two carpels with multiple ovules for Glaucidium – and we get two follicles and many seeds inside.

Let’s count the seeds in this one follicle: you don’t say, precisely enough for 1 pck.!

Glaucidium palmatum fruit and seeds

Call them as you wish

Initially I wrote a short post for Growing Syneilesis from seeds for the germination page, and then I thought, wait a minute, it is not the only Compositae with fruits/seeds related problems.
So, I wrote a longer post on the subject and then thought, who’s going to read all this? Finally, I wrote another shorter post, and there will be more flower pictures later.

I think everyone knows that in Compositae (Asteraceae) what seems to be a single flower is actually a cluster – a composite of smaller flowers (florets), usually called a head which contains disk and/or ray flowers; both types can be bisexual or unisexual.
And, many species require cross-pollination in order to set seeds; the pollen has a special presentation mechanism which prevents self-pollination (I will talk more about this for Syneilesis).

Probably many are also aware that what we call ‘seeds’ in Asteraceae are actually fruits. In simple botanical terms they are single-seeded nuts. Scientifically, they have been called achenes by some and cypselae by others (see note).

So, we don’t see, collect and sow the actual seeds, but the whole fruits.
It is often a deceptive situation because copious amount of fruits are formed but most of them are seedless (cypsela is a very good name to illustrate that) – like it often happens in the case of Syneilesis aconitifolia. Another example, from my garden this time, is Achillea ageratifolia ssp. serbica.

Syneilesis aconitifolia – lots of empty fruits

Achillea ageratifolia ssp. serbica, flowering very well last summer, alas just lots of empty cypsalae were formed

Conclusion:

If we call them fruits or seeds is not that important; I myself call them ‘seeds’. The important thing is to understand the distinction and be able to assess the situation when it comes to sowing, namely if you are really sowing ‘seeds’ or just ‘fluff’ (e.g., empty fruits).

This is why it is always a good idea to carefully check the fruits for many species of this family; take a note on Aster spp. and Solidago, Anaphalis, but not only.
Often is easy to select the good ones (which are plump) when observed with the naked eye; sometimes with the use of a magnifier – and a lot of time…The smaller they are, the harder is to distinguish the good ones.
For large quantities of fruits the cut method can be used by selecting samples.

Anthemis carpatica ssp. pyrethriformis – selected good and empty cypselae using a magnifiyng glass

Sowing seedless fruits, no matter what we call them, and no matter the plant family, will never result in germination.
Read about
Linnaea borealis and Acer triflorum.

Note: Fruits of composites have been called “achenes” because they resemble true achenes. Achenes are dry, hard, single-seeded fruits derived from unicarpellate, superior ovaries. Ovaries of composites are bicarpellate and inferior. Fruits derived from ovaries of composites are called cypselae “ – Flora of North America

cypsela – from the Greek kypselé, a box, hollow vessel


 

 

Well, not quite seedless – here’s one image. I didn’t have time to finish what I wanted to do for the Compositae family and, taking into consideration the weather here, maybe it is for the best. We can really use some colours at this time!

Centaurea salonitana seeds

So, just a gallery with mostly Centaurea and few others thistle-like species :)
The names are displayed on mouse hover, or click and browse through the gallery

 

 

 

I only have a small collection of genera/species from the  Boraginaceae family. I must work better at this family, that’s for sure.
Here they are for now: Onosma, Omphalodes, Pulmonaria and something a bit more rare, Craniospermum.
And because species placed formerly in Hydrophyllaceae are now included in Boraginaceae, there is also Hydrophyllum.

For names, hover over images or click to open the gallery

Only one species/genus is shown, a couple more species are displayed in the new Boraginaceae page.

 

This Friday I indulge in one of my favourite plant families; say it out loud – Berberidaceae!

It is a large family and here I emphasize of course, the ornamental side of it. Many known and beloved garden perennials (usually for the woodland gardens) belong to this family: Epimediums, Vancouveria, Achlys, Podophyllums, Jeffersonia, Ranzania, Diphylleia and so on. From the woody species, Berberis and Mahonia are widely cultivated.

What else they have in common, seeds speaking and referring to the perennials, is the fact that in most cases, these are hydrophilic (i.e. do not tolerate desiccation) and so for optimum results they need to be sown fresh or kept in moist storage.
The pattern required for germination in most cases is WARM/COLD, sometimes with multiple cycles (see Caulophyllum).
Hover over pictures for names

Two genera, Caulophyllum and Gymnospermium, present another rarely seen feature in Angiosperms, namely that the ovary walls burst open and the seeds develop and ripen in a ‘naked’ state (they look like fruits, but nonetheless are just seeds). Fascinating.

For more please see the newly published Berberidaceae page, the 10th in line!

I know seeds are boring (but how else can we obtain plants?), so here are few plant pictures, mostly to show species I don’t have seeds yet: Vancouveria hexandra, Achlys triphylla,   a nice, fat Dysosma and a hybrid Epimedium raised from ‘Amber Queen’ seeds (hybrid). For others like Caulophyllum, Jeffersonia, Podophyllum…I already showed pictures many times and they are also featured in the Seeds shop.

Vancouveria hexandra, garden cultivated; the fruits and seeds are very close with those of Epimedium.

Achlys triphylla, Vanilla leaf (Deer foot) in wild habitat, Victoria Island, BC.

Dysosma versipellis (syn. Podophyllum versipelle) at Butchart Gardens, BC

Epimedium ex. ‘Amber Queen’, a 3-years old plant flowering in my garden (for Epimedium only hybrids will be obtained when growing from seeds)

Only one more picture, more for the purpose to emphasize the name that should be used for Jeffersonia dubia – guilty of charge myself; I’ll try to go the right way from now on.

Plagiorhegma dubium (syn. Jeffersonia dubia)

Additional pictures will be posted later on FB in order to save space here on the website.

 

 

I don’t remember if I’ve already done a Friday’s seeds about Pulsatilla; in any case this is more to draw attention to the Pulsatilla species from the shop.

There are not many, but all are fresh of this year and this is the best time to sow!
See them here in the
Pulsatilla category.

I don’t have many pictures of these gorgeous species, reason why I have to show the seeds. I know they are not as attractive as the flowers but what can I do, and in any case, they can be used for ID purpose.

Pulsatilla albana ssp. armena – the Pulsatilla ‘seeds’ are actually fruits – achenes with “fluffy tails”.

Pulsatilla vernalis


A good idea is to admire pictures while also reading advice on their germination at the same time – all on the Scottish Rock Garden forum! This is the link to Pulsatilla 2013 thread, but there are more topics on Pulsatilla:
http://www.srgc.net/forum/index.php?topic=9988.0

Note: I’ve already sown my batch (I have to practice on this genus) and there are few leftover seeds of Pulsatilla alpina var. alpina, which will come attached as a gift to the first order containing other Pulsatillas.

Pulsatilla alpina ssp. alpina (large, white flowers)

 While talking Ranunculaceae seeds that need to be sown asap, I recommend to also have a look at  Adonis vernalis.

Adonis vernalis

I wanted something special for this arctic day and the woolly seeds of Strophostyles helvola are just perfect; I will stress again the woolly :)

Strophostyles helvola coiled pods and seeds

Strophostyles helvola, trailing wild bean (or amberique bean)  is an annual vine native to eastern Canada and the US; the pea-like flowers are light pink/lilac and they form pods very similar with those of Phaseolus vulgaris; the pod coils when it dries up to release 4-8 woolly  seeds.
The stems will trail on other plants, or if not, will inter-twine with each other on the ground, just like Amphicarpaea bracteata does (google images).

This species has had various medicinal uses for the Native Peoples, and there are also indications that the seeds were used as a food source; they were found in a few archeological sites in NA.

Those interested can read more about this in the curriculum of the Advanced Paleoethnobotany Seminar from Washington University: https://pages.wustl.edu/fritz/strophostyles-helvola-l.-elliot

And, while talking wild foods, it’s winter so new ideas are always welcomed – have a look at this article which recommends other edible species found on the coastal habitat (same where Strophostyles can also be found):

https://ncseagrant.ncsu.edu/coastwatch/previous-issues/2011-2/summer-2011/coastal-wild-edibles-stalking-the-wild-sea-lettuce/

What about: amberique-bean humus with cattails au gratin?
Go foraging this year! :))

A few images recently added to the Caryophyllaceae page on the Seeds Library all species collected this summer in the Carpathian Mts. and Dobrogea region, Romania.
In general, species from Caryophyllaceae family are easy to grow from seeds and some (for example,  think Dianthus, Silene) may even flower in the first year.

Pictures with various Caryophyllaceae showing the habitats they grow in, will follow this weekend. Stay tuned, I’ve prepared many pictures – the time for winter getaways has come!

 

Cerastium alpinum ssp. lanatum- alpine mouse-ear

Gypsophila petraea

 

Silene compacta

 

 

Having too many seeds makes it hard to choose which one to show/write about.
I am going the easy way with the last species uploaded to the shop – the Kalm’s St. John’s wort, a (sub)shrub native of the Great Lakes region in Canada and US.

Hypericum kalmianum seeds – Kalm’s St. John’s wort (Fam. Hypericaceae)

Cultivated to some extent, this Hypericum has narrow bluish-green leaves and cheerful, large golden flowers with extruded stamens in late summer; the capsules mature very late.
It is a showy, adaptable plant; evergreen (or partly) and as one can guess after its wild habitat, with excellent cold hardiness. Another attractive feature is the brown/reddish bark which peels off.

Lots of qualities!

The specific epithet honors Pehr Kalm, one of Linnaeus’ disciples ; an explorer, botanist and naturalist, he traveled to and lived for a while in North America in mid-17th , discovering and writing about plants, animals, insects, Niagara Falls, and in general about the life of American colonies at the time.

Another species that can be found with H. kalmianum, bearing the same name is Lobelia kalmii  shown in the gallery.
Days are short now so you can start to read more about Pehr in the wiki stub and then follow other links: https://en.wikipedia.org/wiki/Pehr_Kalm

Besides the seeds, a sunny image with Echinops crispus (curly Echinops) on this bitterly cold day; the red flowers of Anthyllis might warm us up.

Echinops crispus and Anthyllis vulneraria var. coccinea in the background, early June 2017

I secretly watched this globe thistle evolve from a small seedling into a beautiful clump this year; too bad I was away just when at its most glorious time. The underside of leaves and the stems are covered with a thick pubescence (shown in the featured image), a perfect adaptation for the drought; grown from seeds collected in the southern Ural Mts. (Chelyabinsk region).

Echinops crispus, July 2017

 Like in many other species from Fam. Asteraceae, some of the fruits (achenes) are empty, thus careful selecting of the seeds is necessary.

Echinops crispus fruits and seeds

Echinops name from the Greek words ‘ekhinos’ = hedgehog, sea urchin and
‘ops’ = face, head (referring to the appearance of the inflorescences)

Honestly, I had no intention to write another post so soon, but plants need to be shown at their best moments. And, Amsonia hubrichtii, the Arkansas bluestar is absolutely radiant right now.

Amsonia hubrichtii in late fall

Depending how much sun receives it may also take a russet hue (notice the capsules).

I won’t wonder again about why is not cultivated more; the images speak for themselves. It is a foliage plant by definition, the thread- like leaves will combine beautifully with almost anything else; those who want more flowery display should plant other perennials close to it and let them complement/weave through the delicate foliage.

The flowers are pale blue, not very noticeable but a nice addition.

With Delphinium likiangense; probably larkspurs (Consolida regalis) would make for a nice combination as well.

Without thinking much, one year I planted Dahlia coccinea behind it; now it has become the rule.

The seeds are quite particular, like of other Amsonias (Fam. Apocynaceae); the fruits (capsules) can be spotted in one image above.

Amsonia hubrichtii seeds

PS. It is a bit late to emerge in the spring; planting a bunch of small spring bulbs & daffodils close to the clump will do the trick ;)

 

This fall I collected for the first time fruits of Viburnum acerifolium (maple-leaf Viburnum) and Viburnum rafinesquianum (downy arrow-wood Viburnum).

Surely, I was curious to see how the ‘seeds’ (endocarp + seed) are looking but how about their germination?
There are contradictory statements on the web; some say they first need a cold stratification, others say warm. It’s good to clarify this because I like to provide reliable info.

So, back to the books ;)

Viburnum acerifolium pits (endocarp+ seed)

Viburnum rafinesquianum pits

I like Baskin & Baskin: Seeds – Ecology, Biogeography, and Evolution of Dormancy and Germination (2014) because besides giving results from their own experiments, there are citations of hundreds of other articles on germination (of course, in some cases things are not always clear).

And, the warm stratification is the winner!
These Viburnum spp. have deep simple epicotyl morphophysiological dormancy and the germination requirements are: warm/moist stratification period (during which roots emerge), followed by a cold/moist period (after which the shoots grow).
In translation, if sown in late summer they will show up the next spring; otherwise they will germinate complete in the second spring after sowing.

On the same list are many other Viburnum species like: alnifolium, betulifolium, cassinoides, dentatum, dilatatum, lentago, opulus, parvifolium, prunifolium….Those who don’t have access to the book can read here a very short but conclusive summary regarding Viburnum acerifolium germination:
http://www.bioone.org/doi/full/10.1674/0003-0031%282005%29153%5B0232%3AEDIVAC%5D2.0.CO%3B2

Also another summary here:
https://www.researchgate.net/publication/267996605_Overview_of_seed_dormancy_in_Viburnum_Caprifoliaceae