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Perception

Medicinal plants, Propagation

Vermiculite from Latin ‘vermiculus’ = wormlet

The recently emerged rootlet of bloodroot seed has attached on this vermiculite particle with the same desperation a climber clings onto a rock. A place to grow on, salvation…

Sanguinaria canadensis seedling attached

Sanguinaria canadensis seedling germinated in a bag with vermiculite, growing attached on a vermiculite particle

For us, it remains just an exfoliated fragment of a hydrated silicate mineral; worm-like shaped, lightweight, incombustible, compressible, sterile, with a high cation-exchange capacity…

Only very few Sanguinaria canadensis seeds have started to germinate in moist storage; this one was particularly well developed – good genes probably… The very young rhizome already shows signs of the future red coloration characteristic to Sanguinaria rhizomes.

Sanguinaria canadensis
Sanguinaria canadensis - seedlings

December 9, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/12/Sanguinaria-canadensis-seed-with-rootlet-emerging11.jpg?fit=578%2C341&ssl=1 341 578 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-12-09 18:21:322024-03-26 09:33:34Perception

Check and skotomorphogeneticals

Propagation, Wildflowers of North America

Is this a catchy name, or what?

I got into a routine to check the moist packed seeds at mid and end of the month. Remember the germinated Trilliums? Well, some got planted in pots and some remained in moist vermiculite, which is also a proper medium to easily check on them and take a few more pictures.

You can still read in many places that various Trillium species have a double dormancy but that was really old school thought. Here’s a LINK for something more up to date on what’s been called skotomorphogenetic growth (found in other species too). This concept defines germination as the point when the radicle/rhizome emerges from the seed and all the growth that follows represents the development of the seedling in the dark (from ‘skoto’ – dark in Greek).

Trillium grandiflorum seedling (late November)

Trillium grandiflorum seedling (late November, 2014) -a cute ‘skotomorphogenetic’ with a fatty, little rhizome; the cotyledons are already visible, now it needs more cooling before elongation will start. All this growth was achieved based on the energy reserves stored in the endosperm.

It makes sense. The term double dormancy puts quite a few people off from growing such species from seeds because it implies that they really take a lot of time to germinate. Furthermore, it suggests that the seeds are lying underground and nothing happens, which is not only completely false but also dangerous as you may miss providing the care that they need.

So, skotomorphogenetical it is; I just wish they would have found another name…All the other moist packed seeds are fine; in the featured image – plump, moist seeds of Paris quadrifolia (a Trillium relative, that is also called ‘double dormant’).

Note: one Trillium fruit can have both: seeds with dormant embryos and without – these will start germinating by fall in moist storage (or in pots).

November 27, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/11/Paris-quadrifolia-seeds.jpg?fit=600%2C346&ssl=1 346 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-11-27 09:46:222025-10-21 13:30:23Check and skotomorphogeneticals

The sleeping beauties

Gardens, Propagation

After an early winter arrival I am in the rush now to tuck in safely my sleeping beauties. The garage serves as a ‘winter storage/garden’. In what I call sleeping beauties I include various species done from seeds that have one thing in common: that they are ‘sleeping’ in their first season, totally or partially. Sleeping is not quite a proper term because they are actually growing but mostly underground. However, for a long period of time you are staring and caring for what, some may call, empty pots.

They are mainly hypogeal germinators that in their first season will only emerge the radicle and grow a young tuber/rhizome: Paeonia species, Trilliums, some Lilium species… Paeonia mlokosewitschii seedlings resemble little creatures hungrily sucking water and nutrients from the soil mix ;)

Paeonia mlokosevitschii seedlings
Paeonia and other sleeping beauties
Paeonia rockii - white

Others like Glaucidium palmatum, will germinate totally underground or above, but remain arrested at the cotyledons stage and spend the season fattening up a growing point underground. The whole process can be sped up a bit if you can sow right away in late summer and/or treat with GA3.

 I can already envision their awakening next year, after the Spring’s big kiss!

I know that many don’t care to start from seeds species Paeonies because it will take at least 3-4 years until they start flowering but their spring foliage is equally mesmerizing. For the same reason it is not very easy to find to buy them and when available are quite expensive. Better start them from seed – in a blink of an eye they will grow, prosper and flower! I do not have such seeds to offer yet, but most often you will be able to get hold of a few Paeonia species through Seed Exchanges.

PS. NOT recommended to scratch the pots, or remove the seedlings, like I did; be patient until spring. In my excuse, I needed pictures so I can share the experience.

November 24, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/11/Paeonia-mlokosevitschii1.jpg?fit=600%2C408&ssl=1 408 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-11-24 11:15:422023-03-20 17:41:30The sleeping beauties

No-DOD’s

Propagation, Wildflowers of North America

DOD meaning: Dead on delivery

I don’t know precisely if the DOD term was coined by the renowned Prof. Norman Deno but surely he reminded me of it while reading a delightful Bulletin of the American Rock Garden Society from 1991. Various seeds require to be kept moist after collecting, for storage if not sown right away and for delivery (by Seed companies or in Seed exchanges). Otherwise, they will be actually non-viable when sown later or shipped. There is little to expect of such seeds in terms of germination; hence the DODs. In his article named – “Fatal Treatments of Seed” :) he elaborates on how to kill seeds of 12 species by practicing wrong techniques of storing an germination. 

This is valid for many North American wildflowers, including Trillium species, and among them, Trillium grandiflorum. After collecting (see how the fruit looks when it’s ready), such seeds should be either sown right away or stored moist. When kept at room temperature, some seeds will emerge radicles by fall; then in the spring will send up the first leaves. The rest of the seeds (like in other hypogeal germinating species) will grow a radicle/baby rhizome or tuber in the coming season, and then send up the first leaf only after another cold cycle (that means complete germination in their second year).

These are a few Trillium grandiflorum seeds with emerged radicles that I found last week, during my routine check of the moist stored seeds. A few more pots were added to my collection and I am looking forward to see these little Trilliums sending up their first leaf next year! The other seeds are still in moist storage, awaiting…

Providing moist-packed seeds it is a lot of work but BotanyCa will not be responsible of any DOD’s! See all the Moist-packed Seeds from the Shop.

Trillium grandiflorumAnd here’s the link for the ARGS Bulletin -1991 (open as a pdf). It contains great articles like: Saga of a Woodland Garden by Harold Epstein and Hepaticas and Anemonellas by Jeanie Vesall to mention just a few…

October 27, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/10/Trillium-grandiflorum.jpg?fit=600%2C296&ssl=1 296 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-10-27 09:19:512025-10-21 13:25:43No-DOD’s

The Beauty and the Parthenocarpy – Acer triflorum

Gardens, Plant portraits, Propagation

Beauty is found in almost any maple tree, even if we are to consider only their colourful fall display. The Beast shows its ugly head especially in the case of trifoliate maples and is called parthenocarpy. As a remainder, this means production of seedless fruits (without the fertilization of ovules).

A really undesirable trait, especially for species with indehiscent fruits (samaras, nuts, achene), either if we want their seeds for consumption or to use them for propagation! This is also the explanation for the rarity of some magnificent tree species in our landscapes. Acer griseum, the best known of the trifoliate maples presents parthenocarpy to some extent but not as much as the beauty called Acer triflorum.

Acer triflorum fruit

Acer triflorum

A trifoliate, very hardy maple from Manchuria and Korea “simply an outstanding small specimen maple, lovely foliage, exquisite bark and small habit contribute to the overall landscape effectiveness…; uses for good trees are endless” (Michael Dirr)

I did a little experiment with about the 60 seeds I had and a pruning shear. By nicking the end of the extremely hard samara and then cracking it longitudinally you can extract the seeds, if any. I found 3 viable seeds and 3 aborted (stenospermocarpy) – about 5% viable seeds!

PS. One easy method to separate seedless fruits is to check if they float; the ones with seeds will go to the bottom. This is recommended for many species but actually not necessary; it is useful though for fruits (like samaras and nuts) when it is hard to say if there are any seeds inside.
The fruit wall is so hard in Acer triflorum that it needs lots of stratification just to wear it down, so I preferred to extract the seeds (I didn’t expect to find any, but perseverance paid off :)

October 18, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/10/Acer-triflorum4.jpg?fit=600%2C339&ssl=1 339 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-10-18 12:15:262023-11-09 10:15:19The Beauty and the Parthenocarpy – Acer triflorum

A chic hat

Medicinal plants, Propagation

Update: ID as Aconitum variegatum subsp. paniculatum

All Aconitum species are wearing cool hats but this one in particular with the hood on one side looks very chic. Growing Aconitum from seeds that are not properly identified is quite a pain, but worth the trouble.

All monkshoods are equally beautiful and deadly, with spikes of violet, dark blue, yellow or white hood-shaped, complicated flowers that one needs to know the terminology in order to ‘read’ their characters. Species are usually described on the basis of root and flower morphology.

Aconitum variegatum

Update: Aconitum variegatum subsp. paniculatum

After lots of searches, pictures, dissections… this one was narrowed down to Aconitum variegatum subsp. paniculatum (syn. Aconitum degenii) or A. variegatum subsp. variegatum. We’ll know for sure in late fall after I’ll dig it up and see the tuber shape; and maybe I’ll have a seed or two but glad to hear other opinions…

Aconitum variegatum

Aconitum variegatum: sepals (hood with pronounced rostrum) and petals with coiled nectary spurs

 

Before being praised as a garden plant, Aconitum was considered (and still is) “the king of the poisons” (Europe) or “the king of medicines” (Tibet and China), depending on where it grew, but this is a good subject for wintertime storytelling…
It is a good idea to wear gloves when doing anything that involves touching an Aconitum. It contains highly active and toxic alkaloids.

Aconitum variegatum

Aconitum variegatum (?)

 

September 8, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/09/Aconitum-variegatum-feat..jpg?fit=600%2C352&ssl=1 352 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-09-08 21:06:262023-03-20 17:41:32A chic hat

For Seed trade – Helleborus purpurascens

Botanical travels, Plant portraits, Propagation

Update: since I wrote this post in 2014! we now have the plants grown from wild collected seeds in the Carpathian Mts. (2 different accessions) flowering well and rewarding us with seeds. Nothing is more satisfying than to see plants grown from few seeds found by chance  growing well in the garden. This species is increasingly rare in the wild; growing it in the garden it is also a matter of conservation at this point. The greenish-slate gray with back dusted purple flowers are exquisite! not to mention the foliage.

 

“I am not young enough to know everything” – Oscar Wilde

A quick post because I just ‘discovered’ that Helleborus purpurascens seeds are best sown fresh in the summer. They need to go through a warm-moist period followed by cold-moist in order to germinate best. Older seeds will still germinate but in a lower percentage. Helleborus is one thing I have never done from seed (members of Ranunculaceae are notoriously difficult germinators). This is a rare Helleborus species distributed only along the Carpathian Mountains range, up to central and northern Hungary. Interestingly, it can be found growing in full sun, in open alpine meadows and also under dappled shade in beech woods.

Helleborus purpurascens in native habitatHelleborus purpurascens growing in full sun in native habitat – Carpathian Mts.

Graham Rice, the authority in all Helleborus matters, has an extensive article on H. purpurascens on his website, you can read it HERE. In a bit of a hurry, I will quote him so no one can say I am biased because of its origins:

It is “one of the most captivating species for its engaging habit of flowering so enthusiastically, for its subtle metallic tints… The foliage too is unlike that of other hellebores, in that it radiates from the tip of the petiole in a neat circle.”

As for the flowers, a few forms that are known from Botanical Gardens: “are purple in colour with dark veins, slightly pinkish towards the base and netted towards the edge; some flowers have an overall green haze…In a second form which reaches 15 in in height the flowers are smoky blue-purple in colour, darker outside than in, with slightly reddish veins inside and green nectaries. Other forms may be slate purple or deep purple outside, shining pinkish shades….”

Helleborus purpurascensHelleborus purpurascens – from what’s left it seems matching the description!

A truly collector’s plant, so this is a call for a seeds trade – if someone wishes a few fresh Helleborus purpurascens seeds to sow them right away, please get in touch with me here, on my email or at infoATbotanicallyinclined.org (of course replace AT with @

It would be a pity not to have them all germinating well – still enough summertime left!

Helleborus purpurascens seedsHelleborus purpurascens seeds

 

August 13, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/08/Helleborus-purpurascens-open-fruit.jpg?fit=600%2C383&ssl=1 383 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-08-13 20:10:462023-03-20 17:41:33For Seed trade – Helleborus purpurascens

A risky business – Polygala paucifolia

Plant portraits, Propagation, Wildflowers of North America

I think some are imagining that trying to open a business of selling wild collected seeds is a breeze – happily wandering in fields and mountains and grabbing here and there whatever comes under your eyes. Well, very far away from the truth. For example, who would think about stumbling into a massasauga rattlesnake (the only venomous snake in Ontario), while collecting Polygala paucifolia seeds!

Massasauga snake

Massasauga rattlesnake (Sistrurus catenatus)

The timing to collect Polygala is almost impossible; the fruiting/seed setting is usually low and then the seeds are equipped with elaiosomes that ants will carry away very fast. So, there is no wonder that seeds are almost never offered and, although desirable, neither are the plants! Looking at the flowers, you understand why all this is worthwhile!

Polygala paucifolia

Polygala paucifolia

My germination trials with Polygala seeds showed that dry stored seeds in the fridge, sown in the spring germinate very well, and also the seedlings are developing very well. Seeds stored moist-cold, germinate later and in a lower percentage.

Polygala seedlings

Polygala paucifolia seedlings – germination 95%; they may look small size but remember that this is a little plant

Would you care for some seeds? - click

PS. Keep your eyes wide open when hiking in the rattlesnake habitat! When moving, it makes a low rattle noise to make you aware, but when standing still it is very well camouflaged and it doesn’t rattle. They give birth to live baby-snakes, finger-size but already venomous! It is designated a species at risk in Ontario – more about it on Reptiles and Amphibians of Ontario website.

August 12, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/08/Polygala-paucifolia-capsules-and-seeds.jpg?fit=600%2C311&ssl=1 311 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-08-12 09:41:342023-03-20 17:41:33A risky business – Polygala paucifolia

Rapunzel’s flower – Phyteuma

Alpine plants, Gardens, Plant portraits, Propagation

People in Europe call this member of the bellflower family (Campanulaceae) after many names: devil’s claws (Germany), Oxford Rampion (England), Raponzolo (Italy), and so on. We could definitely give it many other common names; I like to think of it as Rapunzel’s flower.

Phyteuma is strictly a European genus with quite a few species, not very often seen in the gardens. Phyteuma scheuchzeri, flowering now in one of my rock-containers is the most common in cultivation (I was aiming for P. sieberi, maybe next time…).

Phyteuma scheuchzeri

Phyteuma scheuchzeri

It doesn’t look like a bellflower, that’s for sure – it looks much cooler! In Phyteuma species the flowers are grouped in spiked, ball-like inflorescences (aka. floral sea creatures :-) which at full bloom ‘explode’ becoming fluffy. They can be found growing in a variety of habitats, with P. sieberi being the most alpine.

Phyteuma scheuchzeri

Phyteuma scheuchzeri

Another mountain growing Rapunzel’s flower is Phyteuma orbiculare, photographed here in a rich sub-alpine meadow in the Carpathian Mountains:

Phyteuma orbiculare

Phyteuma orbiculare

 Propagation: easy enough from seed (very small, fine seeds just like Campanula, sow superficial); most are warm germinators.

And just because I like word rhymes: Did you know that Phyteuma has a sister named Asyneuma? Another great but very little cultivated member of the bellflower family.

 

June 24, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/06/Phyteuma-scheuchzeri-feat..jpg?fit=600%2C316&ssl=1 316 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-06-24 08:22:342024-06-13 14:37:58Rapunzel’s flower – Phyteuma

In praise of little plants I

Alpine plants, Gardens, Plant portraits, Propagation

Plants that did make sense to have in my small garden

A dwarf, big flowered blue columbine: Aquilegia discolor, most probably a cross (from Seedex as A. saximontana)

Aquilegia discolor (cross)

 Aquilegia discolor cross

True that if we would grow only ‘reasonable’ plants, our gardens would lack all spontaneity and wonder. But because I can now easily enjoy them in containers, and not worry about their relocation, I think a bit of praise is warranted.

On the other side of the container, a tiny hardy ginger: Roscoea tibetica (from Lost Horizons) – very precious, after the bad winter we had, who knows if I will get to see the other Roscoeas from the garden.

Roscoea tibetica

Roscoea tibetica

From another container, the most fragrant, fringed Dianthus I know: Dianthus petraeus (from wild collected seeds in the Carpathian Mts.) Too bad I cannot insert a ‘scratch patch’ with its perfume.

Dianthus petraeus

Dianthus petraeus

A rock jasmine: Androsace sarmentosa – a small piece I saved from an old plant, I hope it will thrive again (or set seeds, or better both).

Androsace sarmentosa

Androsace sarmentosa

and more are on their way to flower…

June 5, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/06/Dianthus-petraeus-featured.jpg?fit=550%2C339&ssl=1 339 550 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-06-05 10:01:402023-03-20 17:41:36In praise of little plants I

A Primula-rina

Gardens, Plant portraits, Propagation
A few weeks ago I stumbled upon this Primula that was showing promise to be a great Primula-rina!
Primula frondosa foliage

The Ballerina – Primula frondosa (a bit elongated in a container in early spring)

Last time when I had visions of dancing plants it was two years ago because of a twirling Arisaema – it is not that often to discover a first class ballerina!.

Primula frondosa is a dwarf, farinose primula endemic from Bulgaria where it grows on cliffs at 900-2000 m altitude, in partly shaded, moist crevices. It is very hardy and will show up from under the snow, with a tight silvery rosette (you may wish that it remains like that), but then the leaves expand and remain powdery only beneath (but the flowering stem and flower pedicels still covered in silvery hairs). A very floriferous primula: umbels with up to 30 pink, delicate flowers with a yellow eye in early spring.

Primula frondosa flowering

Primula frondosa in full bloom right now ( 2-3 seedlings were planted together in the fall)

Sometimes mistaken for P. farinosa (and vice-versa, but P. farinosa has white-farina on both side of the leaves, and it flowers much later). Both are commonly called Bird’s eye Primulas.

Propagation: very easy to grow from seeds (like other Primula spp.), and it will start flowering in the second year – soooo gratifying!

 

May 14, 2014
https://i0.wp.com/botanicallyinclined.org/wp-content/uploads/2014/05/Primula-frondosa-featured.jpg?fit=600%2C339&ssl=1 339 600 BotanyCa https://botanicallyinclined.org/wp-content/uploads/2013/04/logo-red-new2.jpg BotanyCa2014-05-14 09:02:242023-03-20 17:41:58A Primula-rina

The Magic of Germination

Propagation
Using giberellic acid as an aid in germination

More species are germinating and because it’s still cold outside it’s a good time to ‘blag’ a bit about the germination. Each individual seed is a little wonder in itself: it does contain the plant we want – only if we can make it germinate! What I don’t like when growing from seed is not the ‘un-germination’ but the incertitude of what happened – what went wrong? – bad seeds, bad soil mix, too deep, too cold, too dry, not enough light, too much light… Also toooo much information on the web now can make things even more confusing. Here I’ll talk only about what I personally do.

Glaucidium palmatum seed germination

Glaucidium palmatum seed germination

Quite a few species (the most desirable) require ‘special treatments’ for germination like: stratification (moist & cold), alternation of cold and warm periods, incantations, sanding, soaking in GA3, frustrations… you get the idea. If you cannot easily provide a cold and moist period, the treatment with GA3 (acid giberellic) works in some cases wonders. I really like the convenience of GA3, which eliminates some variables from the process.

 Aquilegia-canadensis-semi-double-form-seedlings


Aquilegia canadensis – semi-double flower form seedlings

There is no need to seed way ahead of time or get buried under endless small pots that will get lost in the sway of other spring garden jobs. If the seeds are viable, they’ll germinate; if not, at least you’ll know it wasn’t your ‘brown’ finger at fault. Good to know, however, that GA3 at inappropriate concentrations can also destroy the seeds or lead to poor quality seedlings.

 These are two methods I use:

 1. Keep seeds in their package in the fridge (dry storage). When time to sow, prepare a GA3 solution 500-1000 ppm, soak seeds until next day, plant them in pots, cover with a thin layer of mix – place under lights (or outside if you sow late spring).

 2. For the most recalcitrant – place the seeds into a moist paper towel inside a Ziploc bag, keep in the fridge (moist storage). When time to sow, squeeze the moist paper and add the GA3 solution over seeds, then keep until next day and then sow.

Aquilegia candensis seeds treated
Aquilegia candensis seeds treated
Glaucidium palmatum GA3 treatment
Glaucidium palmatum GA3 treatment
Seeds storage - moist paper in ziploc bags
Seeds storage - moist paper in ziploc bags

 Germination should occur in 1-2(3) weeks.

Most excited about Thalictrums I am trying this year: T. delavayi – a Chinese meadow rue, with large lilac-mauve flowers (petal-like sepals; the one I bought a couple of years ago was really small and didn’t make it) and T. isopyroides with a really tiny, steel blue foliage that can grow in full sun – would be good for a rockery (from Turkey, Iran, Syria, Altai Mts.).

Thalictrum delavayi seedlings (GA3 treatment)
Thalictrum delavayi seedlings (GA3 treatment)
Thalictrum delavayi flowers
Thalictrum delavayi flowers

Note: Aquilegia also prefer/require light for germination so you should cover them with a very fine layer of potting mix (in case you forget about this they’ll still germinate but much slower).

 

 

 

 

March 28, 2014
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