Category: Propagation - Page 32 - Botanically Inclined Plants Seeds

No-DOD’s

Propagation, Wildflowers of North America

DOD meaning: Dead on delivery

I don’t know precisely if the DOD term was coined by the renowned Prof. Norman Deno but surely he reminded me of it while reading a delightful Bulletin of the American Rock Garden Society from 1991. Various seeds require to be kept moist after collecting, for storage if not sown right away and for delivery (by Seed companies or in Seed exchanges). Otherwise, they will be actually non-viable when sown later or shipped. There is little to expect of such seeds in terms of germination; hence the DODs. In his article named – “Fatal Treatments of Seed” :) he elaborates on how to kill seeds of 12 species by practicing wrong techniques of storing an germination.

This is valid for many North American wildflowers, including Trillium species, and among them, Trillium grandiflorum. After collecting (see how the fruit looks when it’s ready), such seeds should be either sown right away or stored moist. When kept at room temperature, some seeds will emerge radicles by fall; then in the spring will send up the first leaves. The rest of the seeds (like in other hypogeal germinating species) will grow a radicle/baby rhizome or tuber in the coming season, and then send up the first leaf only after another cold cycle (that means complete germination in their second year).

These are a few Trillium grandiflorum seeds with emerged radicles that I found last week, during my routine check of the moist stored seeds. A few more pots were added to my collection and I am looking forward to see these little Trilliums sending up their first leaf next year! The other seeds are still in moist storage, awaiting…

Providing moist-packed seeds it is a lot of work but BotanyCa will not be responsible of any DOD’s! See all the Moist-packed Seeds from the Shop.

And here’s the link for the ARGS Bulletin -1991 (open as a pdf). It contains great articles like: Saga of a Woodland Garden by Harold Epstein and Hepaticas and Anemonellas by Jeanie Vesall to mention just a few…

October 27, 2014

The Beauty and the Parthenocarpy – Acer triflorum

Gardens, Plant portraits, Propagation

Beauty is found in almost any maple tree, even if we are to consider only their colourful fall display. The Beast shows its ugly head especially in the case of trifoliate maples and is called parthenocarpy. As a remainder, this means production of seedless fruits (without the fertilization of ovules).

A really undesirable trait, especially for species with indehiscent fruits (samaras, nuts, achene), either if we want their seeds for consumption or to use them for propagation! This is also the explanation for the rarity of some magnificent tree species in our landscapes. Acer griseum, the best known of the trifoliate maples presents parthenocarpy to some extent but not as much as the beauty called Acer triflorum.

Acer triflorum

A trifoliate, very hardy maple from Manchuria and Korea “simply an outstanding small specimen maple, lovely foliage, exquisite bark and small habit contribute to the overall landscape effectiveness…; uses for good trees are endless” (Michael Dirr)

I did a little experiment with about the 60 seeds I had and a pruning shear. By nicking the end of the extremely hard samara and then cracking it longitudinally you can extract the seeds, if any. I found 3 viable seeds and 3 aborted (stenospermocarpy) – about 5% viable seeds!

PS. One easy method to separate seedless fruits is to check if they float; the ones with seeds will go to the bottom. This is recommended for many species but actually not necessary; it is useful though for fruits (like samaras and nuts) when it is hard to say if there are any seeds inside.
The fruit wall is so hard in Acer triflorum that it needs lots of stratification just to wear it down, so I preferred to extract the seeds (I didn’t expect to find any, but perseverance paid off :)

October 18, 2014

A chic hat

Medicinal plants, Propagation

Update: ID as Aconitum variegatum subsp. paniculatum

All Aconitum species are wearing cool hats but this one in particular with the hood on one side looks very chic. Growing Aconitum from seeds that are not properly identified is quite a pain, but worth the trouble.

All monkshoods are equally beautiful and deadly, with spikes of violet, dark blue, yellow or white hood-shaped, complicated flowers that one needs to know the terminology in order to ‘read’ their characters. Species are usually described on the basis of root and flower morphology.

Update: Aconitum variegatum subsp. paniculatum

After lots of searches, pictures, dissections… this one was narrowed down to Aconitum variegatum subsp. paniculatum (syn. Aconitum degenii) or A. variegatum subsp. variegatum. We’ll know for sure in late fall after I’ll dig it up and see the tuber shape; and maybe I’ll have a seed or two but glad to hear other opinions…

Aconitum variegatum: sepals (hood with pronounced rostrum) and petals with coiled nectary spurs

Before being praised as a garden plant, Aconitum was considered (and still is) “the king of the poisons” (Europe) or “the king of medicines” (Tibet and China), depending on where it grew, but this is a good subject for wintertime storytelling…
It is a good idea to wear gloves when doing anything that involves touching an Aconitum. It contains highly active and toxic alkaloids.

Aconitum variegatum (?)

September 8, 2014

For Seed trade – Helleborus purpurascens

Botanical travels, Plant portraits, Propagation

Update: since I wrote this post in 2014! we now have the plants grown from wild collected seeds in the Carpathian Mts. (2 different accessions) flowering well and rewarding us with seeds. Nothing is more satisfying than to see plants grown from few seeds found by chance growing well in the garden. This species is increasingly rare in the wild; growing it in the garden it is also a matter of conservation at this point. The greenish-slate gray with back dusted purple flowers are exquisite! not to mention the foliage.

“I am not young enough to know everything” – Oscar Wilde

A quick post because I just ‘discovered’ that Helleborus purpurascens seeds are best sown fresh in the summer. They need to go through a warm-moist period followed by cold-moist in order to germinate best. Older seeds will still germinate but in a lower percentage. Helleborus is one thing I have never done from seed (members of Ranunculaceae are notoriously difficult germinators). This is a rare Helleborus species distributed only along the Carpathian Mountains range, up to central and northern Hungary. Interestingly, it can be found growing in full sun, in open alpine meadows and also under dappled shade in beech woods.

Helleborus purpurascens growing in full sun in native habitat – Carpathian Mts.

Graham Rice, the authority in all Helleborus matters, has an extensive article on H. purpurascens on his website, you can read it HERE. In a bit of a hurry, I will quote him so no one can say I am biased because of its origins:

It is “one of the most captivating species for its engaging habit of flowering so enthusiastically, for its subtle metallic tints… The foliage too is unlike that of other hellebores, in that it radiates from the tip of the petiole in a neat circle.”

As for the flowers, a few forms that are known from Botanical Gardens: “are purple in colour with dark veins, slightly pinkish towards the base and netted towards the edge; some flowers have an overall green haze…In a second form which reaches 15 in in height the flowers are smoky blue-purple in colour, darker outside than in, with slightly reddish veins inside and green nectaries. Other forms may be slate purple or deep purple outside, shining pinkish shades….”

Helleborus purpurascens – from what’s left it seems matching the description!

A truly collector’s plant, so this is a call for a seeds trade – if someone wishes a few fresh Helleborus purpurascens seeds to sow them right away, please get in touch with me here, on my email or at infoATbotanicallyinclined.org (of course replace AT with @

It would be a pity not to have them all germinating well – still enough summertime left!

Helleborus purpurascens seeds

August 13, 2014

A risky business – Polygala paucifolia

Plant portraits, Propagation, Wildflowers of North America

I think some are imagining that trying to open a business of selling wild collected seeds is a breeze – happily wandering in fields and mountains and grabbing here and there whatever comes under your eyes. Well, very far away from the truth. For example, who would think about stumbling into a massasauga rattlesnake (the only venomous snake in Ontario), while collecting Polygala paucifolia seeds!

: Massasauga rattlesnake (Sistrurus catenatus)

The timing to collect Polygala is almost impossible; the fruiting/seed setting is usually low and then the seeds are equipped with elaiosomes that ants will carry away very fast. So, there is no wonder that seeds are almost never offered and, although desirable, neither are the plants! Looking at the flowers, you understand why all this is worthwhile!

: Polygala paucifolia

My germination trials with Polygala seeds showed that dry stored seeds in the fridge, sown in the spring germinate very well, and also the seedlings are developing very well. Seeds stored moist-cold, germinate later and in a lower percentage.

: Polygala paucifolia seedlings – germination 95%; they may look small size but remember that this is a little plant

Would you care for some seeds? - click

PS. Keep your eyes wide open when hiking in the rattlesnake habitat! When moving, it makes a low rattle noise to make you aware, but when standing still it is very well camouflaged and it doesn’t rattle. They give birth to live baby-snakes, finger-size but already venomous! It is designated a species at risk in Ontario – more about it on Reptiles and Amphibians of Ontario website.

August 12, 2014

Rapunzel’s flower – Phyteuma

Alpine plants, Gardens, Plant portraits, Propagation

People in Europe call this member of the bellflower family (Campanulaceae) after many names: devil’s claws (Germany), Oxford Rampion (England), Raponzolo (Italy), and so on. We could definitely give it many other common names; I like to think of it as Rapunzel’s flower.

Phyteuma is strictly a European genus with quite a few species, not very often seen in the gardens. Phyteuma scheuchzeri, flowering now in one of my rock-containers is the most common in cultivation (I was aiming for P. sieberi, maybe next time…).

Phyteuma scheuchzeri

It doesn’t look like a bellflower, that’s for sure – it looks much cooler! In Phyteuma species the flowers are grouped in spiked, ball-like inflorescences (aka. floral sea creatures :-) which at full bloom ‘explode’ becoming fluffy. They can be found growing in a variety of habitats, with P. sieberi being the most alpine.

Phyteuma scheuchzeri

Another mountain growing Rapunzel’s flower is Phyteuma orbiculare, photographed here in a rich sub-alpine meadow in the Carpathian Mountains:

Phyteuma orbiculare

Propagation: easy enough from seed (very small, fine seeds just like Campanula, sow superficial); most are warm germinators.

And just because I like word rhymes: Did you know that Phyteuma has a sister named Asyneuma? Another great but very little cultivated member of the bellflower family.

June 24, 2014

In praise of little plants I

Alpine plants, Gardens, Plant portraits, Propagation

Plants that did make sense to have in my small garden

A dwarf, big flowered blue columbine: Aquilegia discolor, most probably a cross (from Seedex as A. saximontana)

: Aquilegia discolor cross

True that if we would grow only ‘reasonable’ plants, our gardens would lack all spontaneity and wonder. But because I can now easily enjoy them in containers, and not worry about their relocation, I think a bit of praise is warranted.

On the other side of the container, a tiny hardy ginger: Roscoea tibetica (from Lost Horizons) – very precious, after the bad winter we had, who knows if I will get to see the other Roscoeas from the garden.

: Roscoea tibetica

From another container, the most fragrant, fringed Dianthus I know: Dianthus petraeus (from wild collected seeds in the Carpathian Mts.) Too bad I cannot insert a ‘scratch patch’ with its perfume.

: Dianthus petraeus

A rock jasmine: Androsace sarmentosa – a small piece I saved from an old plant, I hope it will thrive again (or set seeds, or better both).

: Androsace sarmentosa

and more are on their way to flower…

June 5, 2014

A Primula-rina

Gardens, Plant portraits, Propagation

A few weeks ago I stumbled upon this Primula that was showing promise to be a great Primula-rina!

The Ballerina – Primula frondosa (a bit elongated in a container in early spring)

Last time when I had visions of dancing plants it was two years ago because of a twirling Arisaema – it is not that often to discover a first class ballerina!.

Primula frondosa is a dwarf, farinose primula endemic from Bulgaria where it grows on cliffs at 900-2000 m altitude, in partly shaded, moist crevices. It is very hardy and will show up from under the snow, with a tight silvery rosette (you may wish that it remains like that), but then the leaves expand and remain powdery only beneath (but the flowering stem and flower pedicels still covered in silvery hairs). A very floriferous primula: umbels with up to 30 pink, delicate flowers with a yellow eye in early spring.

Primula frondosa in full bloom right now ( 2-3 seedlings were planted together in the fall)

Sometimes mistaken for P. farinosa (and vice-versa, but P. farinosa has white-farina on both side of the leaves, and it flowers much later). Both are commonly called Bird’s eye Primulas.

Propagation: very easy to grow from seeds (like other Primula spp.), and it will start flowering in the second year – soooo gratifying!

May 14, 2014

The Magic of Germination

Propagation

Using giberellic acid as an aid in germination

More species are germinating and because it’s still cold outside it’s a good time to ‘blag’ a bit about the germination. Each individual seed is a little wonder in itself: it does contain the plant we want – only if we can make it germinate! What I don’t like when growing from seed is not the ‘un-germination’ but the incertitude of what happened – what went wrong? – bad seeds, bad soil mix, too deep, too cold, too dry, not enough light, too much light… Also toooo much information on the web now can make things even more confusing. Here I’ll talk only about what I personally do.

Glaucidium palmatum seed germination

Quite a few species (the most desirable) require ‘special treatments’ for germination like: stratification (moist & cold), alternation of cold and warm periods, incantations, sanding, soaking in GA3, frustrations… you get the idea. If you cannot easily provide a cold and moist period, the treatment with GA3 (acid giberellic) works in some cases wonders. I really like the convenience of GA3, which eliminates some variables from the process.

Aquilegia-canadensis-semi-double-form-seedlings

Aquilegia canadensis – semi-double flower form seedlings

There is no need to seed way ahead of time or get buried under endless small pots that will get lost in the sway of other spring garden jobs. If the seeds are viable, they’ll germinate; if not, at least you’ll know it wasn’t your ‘brown’ finger at fault. Good to know, however, that GA3 at inappropriate concentrations can also destroy the seeds or lead to poor quality seedlings.

These are two methods I use:

1. Keep seeds in their package in the fridge (dry storage). When time to sow, prepare a GA3 solution 500-1000 ppm, soak seeds until next day, plant them in pots, cover with a thin layer of mix – place under lights (or outside if you sow late spring).

2. For the most recalcitrant – place the seeds into a moist paper towel inside a Ziploc bag, keep in the fridge (moist storage). When time to sow, squeeze the moist paper and add the GA3 solution over seeds, then keep until next day and then sow.

Germination should occur in 1-2(3) weeks.

Most excited about Thalictrums I am trying this year: T. delavayi – a Chinese meadow rue, with large lilac-mauve flowers (petal-like sepals; the one I bought a couple of years ago was really small and didn’t make it) and T. isopyroides with a really tiny, steel blue foliage that can grow in full sun – would be good for a rockery (from Turkey, Iran, Syria, Altai Mts.).

Note: Aquilegia also prefer/require light for germination so you should cover them with a very fine layer of potting mix (in case you forget about this they’ll still germinate but much slower).

March 28, 2014

A study in contrast – Aquilegia species

Plant portraits, Propagation

A group of plants that I really like and hope to increase my collection, are the columbines: Aquilegia spp., and in particular, of course, the alpine columbines. In contrast with the more regular garden Aquilegia varieties, the alpine ones are short in stature but bearing large flowers. In most cases they have a delightful bluish, compact foliage, which in itself makes a wonderful addition to any small rock garden.

Aquilegia scopulorum x coerulea in the rock garden at Wrightman Alpines

All Aquilegia are important food source plants for bees, bumblebees, hummingbirds and hawk moths. Even just for this reason one should include them in the garden. Interesting fact, biologists found that the length of the nectar spurs in Aquilegia evolved to allow flowers to match the tongue lengths of their pollinators. Species with very long nectar spurs, like A. coerulea are pollinated by hummingbirds and hawk moths, while the short spured species, like A. canadensis are fancied by bees and other short tongue insects.

Aquilegia coerulea (Colorado blue columbine) has flowers with very long nectar spurs that look like space ships or sea creatures, wherever your imagination tends to go, up to the sky or down in the ocean. Very variable in height, anywhere from 15 to 90 cm and in flower colours – from white to pale or dark blue; also there are reports of a variety with spurless flowers!

I am greedy when it comes to columbines. This spring I am looking forward to see flowering (and take more pictures) in my rockery a few alpine columbines I grew from seeds: A. jonesii – the smallest of columbines, A. saximontana and A. discolor (that is if my seedlings survive the record low temperatures we have this winter).

Also ready to greet the pollinators this year: A. nigricans (Carpathians Mts. collection), A. alpina and A. atrata.

I would note that most Aquilegia species are polymorphic and difficult to define adequately. Some of the variability is because of introgressive hybridization (Flora of NA). Even distantly related species of columbines are often freely inter-fertile, hence the multitude of hybrids and cultivated forms available. Also, this poses a problem for seeds collection, especially in the case of cultivated varieties if one requires true to type species.

February 26, 2014

In a mood for Arisaema part II – Arisaema galeatum

Gardens, Plant portraits, Propagation

Another great Arisaema that flowers in early spring is Arisaema galeatum. It is another story than A. sazensoo because it has a really big tuber. It is said that can grow to half a kilogram! Last year when I was checking the tuber in early March I caught it just starting to grow and it looked very appealing to me – with a dark-chocolate coating and raspberry syrup on top would be delicious! But I put myself together… This is an Arisaema from the Himalayan range (NE India to Bhutan), which grows during the mansoon months and then the leaves start withering in late summer and goes dormant early. Good to keep this in mind as it shows its requirements for a very good drainage from late summer to fall, and during winter of course.

The flower emerge on the same time with the leaves, on a short peduncle and it has a helmet-like (galeate) spathe, similar with A. ringens. It can be green or brown with whitish veins and has a white, translucent spadix that ends in a thread like whip. I like to call it Dolphin cobra lily, because that’s what it suggested to me first time when it bloomed – a dolphin emerging for air from within the leaf!

Just like its sister, A. ringens, the huge trifoliolate leaf is very ornamental. Actually, I consider it among the most beautiful from all Arisaema species I’ve seen. It unfolds slowly and the back pattern with accentuated purple ribs makes it mesmerizing to watch.

It did form two tuberlets two years ago (not a great rate of offsetting), from which one even produced a small flower in its first season! Unfortunately, the rainy weather we had late summer to fall it proved fatal for the smaller size tubers. If someone wants to give it a try I suggest container culture, so it can be moved to a dry place in late summer, or if in the garden a real well drained area, like close to a tree or shrub that would remove the excess water and also provide the part-shade required.

February 25, 2014

In a mood for Arisaema – Arisaema sazensoo

Gardens, Plant portraits, Propagation

This is an updated post on Arisaema sazensoo – I have more ‘data’ to share now than last year. I am always in a mood for any Arisaema, but especially for the rare ones like A. sazensoo.

Arisaema sazensoo, is one of the first Arisaema to emerge in the spring, just like its cousin A. sikokianum. It is native from Kyushu, Japan and resemble a little A. sikokianum but the spadix doesn’t have such a pronounced white ‘pestle’. The spathe is usually deep purple, recurved over the spadix and the leaves are trifoliolate, like you can see in the images. It was thought to resemble a Buddhist monk in meditation – ‘zazen’, hence its name sazensoo, or at least that’s what I read. Anyway, you can tell it is a very charismatic Arisaema!

Arisaema sazensoo

Another characteristic is that it stays in flower over a very long period of time, comparing with other Arisaemas. It had one attempt to form seeds, which proved sterile, but two years ago in late fall I had the very pleasant surprise to find that it had produced an offset (a tuberlet)!

On a few websites you’ll read that A. sazensoo is a non-offsetting species, but obviously someone got it wrong. In the images below I can present now the tuberlet that has grown quite well in one season (A. sazensoo doesn’t have a big size flowering tuber). More than this, the old tuber shows very clear another tuberlet (which is best left to detach by itself).

Like many other Arisaema species, it prefers a part-shade location and can be grown very well in a container, where a good drainage can be easily provided. Best transplanted in late fall with fresh potting mix and kept dry over the winter.

February 24, 2014