
Paeonia mlokosewitchii shoots after cold stratification; the first stage, roots growing was shown in the previous post (Cold stratification: part 3, the practicals)
Finally the spring arrives – only in our story about the cold stratification 😊We tried our best sowing and providing the cold stratification.
Is there any guarantee that all seeds will germinate well?
Of course not, hope is good, but kept on the realistic side. Sowing and germinating seeds is not a precise science, and many factors, some out of our control, can contribute to a low germination or no germination at all; reason why keeping your pots for at least another year it is always a good idea.
However, at least some species will germinate, and then you may have to deal with too many seedlings! I always recommend to sow smaller batches of seeds every year, instead of 30 pots at once. And, combine easy germinating species with some of the more difficult ones.
What to do when the weather starts to warm up?
I would like to say that we just sit and watch the seeds germinating, but it’s not quite like that 😊

Caulophyllum thalictroides first shoots, after direct sowing (2 years germinator!); work carefully in the spring around the spots where you’ve sown seeds in the ground.
Usually a visible warm up may happen in March in our region, sometimes even later. Not all winters are the same. Unlike the groundhogs, I predict that we will have a very late spring, with short interludes of warm weather here and there.
In case of direct sowing, after the snowmelt, remove the layer of dead leaves/mulch or any other materials you used for extra protection. The best is still to come weather wise, so watch the spot for germination without scratching the ground! In shaded locations it is a good idea to sprinkle a bit of slug pellets around (for just in case).
In case of sowing in moss, there is nothing else to do than waiting for the germination. If too many seedlings appear you will have to transplant some of them later in the season.
In case you sowed in pots, after the snow has melted, open the box(es), or remove any other wrappings from the trays and cold frame to inspect the pots. Most of them will be frozen. Do not worry (follow cold stratification parts 1,2,3); slowly they will start to warm up.
Ideally, boxes and flats with pots should be lifted from the ground on a higher surface to receive more light = warmth. It also makes it easier to keep an eye on them. If March is still very cold, then do this in April.
In case of using a cold frame: open the lid as soon as possible and remove some of the protecting materials. Make a thorough inspection for slugs (eggs can also be at the bottom of pots). Let the lid open as much as possible during warm, sunny days and close it back at night. Most likely you can let go of a part of the wrapping layers.
There is not much to do in the garden so early in the spring. To fuss around the cold frame is a very nice activity 😊 not to mention that early germinating species can take you by surprise.
With the pots buried in the ground there is usually more work to be done as soon as the soil warms up, in case the location is not proper for germination (for example a heavy shaded area). Larger containers are usually to be kept in place; remove the layer of leaves/mulch if it was added and check for slugs; in recent years also cutworms have become a real problem (and they are active at very low temperatures).
If the location is not proper for germination, take the pots out, organize them in a tray/box and lift them up on an elevated surface.
Leaving them buried in the ground, will either delay the germination or expose the fresh, succulent seedlings to various ‘accidents’. Check the bottom of these pots for little slugs or slug eggs; often they hide in between the pots in late fall waiting hungrily for the first ‘victims’. The emerging seedlings can be devoured so fast that it may seem there was no germination at all!
Some species are germinating at low temperatures regardless of the cold stratification method. Depending on the temperatures the first signs of germination are possible in March in our climate even when the pots are looking somewhat ‘frozen’.
From my experience, among the early spring germinators are: Adonis, Corydalis species, Claytonia, Fritillaria, Galanthus, Crocus species, Erythroniums, Hepatica, Helleborus, some species peonies.
How to handle our pots, without a cold frame
If you provided cold stratification underneath your patio table/bench, it is very simple, lift them up. It is not that they can be used for something else until May at least.
A
good, not expensive investment it is a small, plastic ‘greenhouse’ with a detachable cover. It is light and can be moved around; when it really warms up, replace the plastic cover with a shade cloth.
I also use a folding plastic table and I recommend it as a good option.
Advantages: they come in various sizes, are easy to move around to the best locations from early spring to summer, you can put the flats underneath when it rains heavily…also they have a very good work-height for transplanting the seedlings later! When you’re done, fold it back and put it away.
During the spring (meaning late May!), be on guard for heavy rains, late frosts and not to mention late flurries!!! The pots and the emerging seedlings need protection. Keep some plastic sheets/fleece blankets handy.
If you followed the fridge cold/moist stratification, you need to constantly check for signs of germination. Most times not all seeds germinate at once. Wait until a good percentage of seeds have developed radicles and gently plant them in a pot and keep them outdoors if weather allows or at room temperature (it depends when it happens, so you may need indoor lights).
Will all seeds germinate at once?
Many species have a uniform germination. So, after the seedlings grow up a bit, they can be pricked out, if necessary. * Not all species require pricking out in the first season.
However, other species may germinate gradually over the spring, or over a couple of years (Delphinium fissum, Hepatica, Sanguinaria sometimes, etc.).
In this cases, we believe to be ‘poor germination’, when in fact it is the plant’s mechanism of germinating gradually to give its seedlings better chances of surviving. Wait until you have a few good seedlings to transplant or, keep the pot ‘as is’ until next year.
What happens if species that are actually warm germinators were sown in the fall/winter?
Nothing bad; good seeds should germinate in late spring with the raise of temperature, if they were sown accordingly with their germination requirements.
Before using an indoor lighting system which allows for an early sowing, all species were sown in late fall/early winter and all were overwintered outdoors, either under the snow or in the cold frame. It was never a problem for the usual warm germinating species. Cirsium cannum was overwintered in a tray under snow (shown in the previous post). Other well known warm germinators like Centaurea and Gypsophila species were also sown in late fall with good results.
What to do with the non-germinated pots?
As I mentioned, some species are early germinators at lower temperatures, meaning late February to March, depending on the weather.
From my experience, most other species will germinate between April to early May, in our region. There may be exceptions (Disocorea caucasica for example) but generally speaking if something had not germinated by mid June it won’t happen later in the season.
Organize these pots and keep them together in a part-shaded spot, or again in a box; redo the labeling. Do not forget to check the moisture throughout the season. By late fall, repeat the cold stratification. Some of the very early spring germinators (like Corydalis, Claytonia, Crocus) become dormant in June, so you can group all these pots together.
As I tried to explain, there are many reasons for failed germination. Sometimes, seemingly good looking seeds may have internal defects impeding embryo development or germination. When we sow the actual fruits, we cannot even have a visual assessment of the actual seeds.
Some species naturally need repeated cycles of warm/cold, meaning 2-3 years until germination. Not many people are keeping and taking good care of their pots for so long.
However, sowing every year a few pots with a variety of species it is the key of having seeds germinating, seedlings growing and new plants flowering every season!
*Special attention to all species from Aster family: most of them are warm germinators. No matter how you sowed them, they should germinate in the spring. If they don’t, most likely the fruits/seeds were not good: Asteraceae: seeds, achenes and cypsaele
The Germination page has been reorganized with the cold stratification subject listed under Treatments for seeds germination.
Cold stratification: part 1, dormancy
Cold stratification: part 2, FAQ
Cold stratification: part 3, the practicals
Cold stratification: part 4, springtime
PropagationPaeonia mlokosewitchii shoots after cold stratification; the first stage, roots growing was shown in the previous post (Cold stratification: part 3, the practicals)
Finally the spring arrives – only in our story about the cold stratification 😊We tried our best sowing and providing the cold stratification.
Is there any guarantee that all seeds will germinate well?
Of course not, hope is good, but kept on the realistic side. Sowing and germinating seeds is not a precise science, and many factors, some out of our control, can contribute to a low germination or no germination at all; reason why keeping your pots for at least another year it is always a good idea.
However, at least some species will germinate, and then you may have to deal with too many seedlings! I always recommend to sow smaller batches of seeds every year, instead of 30 pots at once. And, combine easy germinating species with some of the more difficult ones.
What to do when the weather starts to warm up?
I would like to say that we just sit and watch the seeds germinating, but it’s not quite like that 😊
Caulophyllum thalictroides first shoots, after direct sowing (2 years germinator!); work carefully in the spring around the spots where you’ve sown seeds in the ground.
Usually a visible warm up may happen in March in our region, sometimes even later. Not all winters are the same. Unlike the groundhogs, I predict that we will have a very late spring, with short interludes of warm weather here and there.
In case of direct sowing, after the snowmelt, remove the layer of dead leaves/mulch or any other materials you used for extra protection. The best is still to come weather wise, so watch the spot for germination without scratching the ground! In shaded locations it is a good idea to sprinkle a bit of slug pellets around (for just in case).
In case of sowing in moss, there is nothing else to do than waiting for the germination. If too many seedlings appear you will have to transplant some of them later in the season.
In case you sowed in pots, after the snow has melted, open the box(es), or remove any other wrappings from the trays and cold frame to inspect the pots. Most of them will be frozen. Do not worry (follow cold stratification parts 1,2,3); slowly they will start to warm up.
Ideally, boxes and flats with pots should be lifted from the ground on a higher surface to receive more light = warmth. It also makes it easier to keep an eye on them. If March is still very cold, then do this in April.
In case of using a cold frame: open the lid as soon as possible and remove some of the protecting materials. Make a thorough inspection for slugs (eggs can also be at the bottom of pots). Let the lid open as much as possible during warm, sunny days and close it back at night. Most likely you can let go of a part of the wrapping layers.
There is not much to do in the garden so early in the spring. To fuss around the cold frame is a very nice activity 😊 not to mention that early germinating species can take you by surprise.
Corydalis seedlings, frame April 30 2019
Claytonia virginica, March 2018
Galanthus, cold frame 2018
With the pots buried in the ground there is usually more work to be done as soon as the soil warms up, in case the location is not proper for germination (for example a heavy shaded area). Larger containers are usually to be kept in place; remove the layer of leaves/mulch if it was added and check for slugs; in recent years also cutworms have become a real problem (and they are active at very low temperatures).
If the location is not proper for germination, take the pots out, organize them in a tray/box and lift them up on an elevated surface.
Leaving them buried in the ground, will either delay the germination or expose the fresh, succulent seedlings to various ‘accidents’. Check the bottom of these pots for little slugs or slug eggs; often they hide in between the pots in late fall waiting hungrily for the first ‘victims’. The emerging seedlings can be devoured so fast that it may seem there was no germination at all!
Anemone quinquefolia, in ground stratification 2018
Container sowing, ground 2018
Corydalis-solida-seedlings, in ground stratification 2017
Some species are germinating at low temperatures regardless of the cold stratification method. Depending on the temperatures the first signs of germination are possible in March in our climate even when the pots are looking somewhat ‘frozen’.
From my experience, among the early spring germinators are: Adonis, Corydalis species, Claytonia, Fritillaria, Galanthus, Crocus species, Erythroniums, Hepatica, Helleborus, some species peonies.
How to handle our pots, without a cold frame
If you provided cold stratification underneath your patio table/bench, it is very simple, lift them up. It is not that they can be used for something else until May at least.
good, not expensive investment it is a small, plastic ‘greenhouse’ with a detachable cover. It is light and can be moved around; when it really warms up, replace the plastic cover with a shade cloth.
A
I also use a folding plastic table and I recommend it as a good option.
Advantages: they come in various sizes, are easy to move around to the best locations from early spring to summer, you can put the flats underneath when it rains heavily…also they have a very good work-height for transplanting the seedlings later! When you’re done, fold it back and put it away.
During the spring (meaning late May!), be on guard for heavy rains, late frosts and not to mention late flurries!!! The pots and the emerging seedlings need protection. Keep some plastic sheets/fleece blankets handy.
If you followed the fridge cold/moist stratification, you need to constantly check for signs of germination. Most times not all seeds germinate at once. Wait until a good percentage of seeds have developed radicles and gently plant them in a pot and keep them outdoors if weather allows or at room temperature (it depends when it happens, so you may need indoor lights).
Will all seeds germinate at once?
Many species have a uniform germination. So, after the seedlings grow up a bit, they can be pricked out, if necessary. * Not all species require pricking out in the first season.
However, other species may germinate gradually over the spring, or over a couple of years (Delphinium fissum, Hepatica, Sanguinaria sometimes, etc.).
In this cases, we believe to be ‘poor germination’, when in fact it is the plant’s mechanism of germinating gradually to give its seedlings better chances of surviving. Wait until you have a few good seedlings to transplant or, keep the pot ‘as is’ until next year.
Nothing bad; good seeds should germinate in late spring with the raise of temperature, if they were sown accordingly with their germination requirements.
Before using an indoor lighting system which allows for an early sowing, all species were sown in late fall/early winter and all were overwintered outdoors, either under the snow or in the cold frame. It was never a problem for the usual warm germinating species. Cirsium cannum was overwintered in a tray under snow (shown in the previous post). Other well known warm germinators like Centaurea and Gypsophila species were also sown in late fall with good results.
What to do with the non-germinated pots?
As I mentioned, some species are early germinators at lower temperatures, meaning late February to March, depending on the weather.
From my experience, most other species will germinate between April to early May, in our region. There may be exceptions (Disocorea caucasica for example) but generally speaking if something had not germinated by mid June it won’t happen later in the season.
Organize these pots and keep them together in a part-shaded spot, or again in a box; redo the labeling. Do not forget to check the moisture throughout the season. By late fall, repeat the cold stratification. Some of the very early spring germinators (like Corydalis, Claytonia, Crocus) become dormant in June, so you can group all these pots together.
As I tried to explain, there are many reasons for failed germination. Sometimes, seemingly good looking seeds may have internal defects impeding embryo development or germination. When we sow the actual fruits, we cannot even have a visual assessment of the actual seeds.
Some species naturally need repeated cycles of warm/cold, meaning 2-3 years until germination. Not many people are keeping and taking good care of their pots for so long.
However, sowing every year a few pots with a variety of species it is the key of having seeds germinating, seedlings growing and new plants flowering every season!
Saxifraga cuneifolia ssp. robusta, cold stratification in moss
Helleborus caucasicus seedlings, 2018
Thalictrum thalictroides
Zigadenus elegans ssp. glaucus seedlings
Geum triflorum seedlings
Gillenia trifoliata, 2016
*Special attention to all species from Aster family: most of them are warm germinators. No matter how you sowed them, they should germinate in the spring. If they don’t, most likely the fruits/seeds were not good: Asteraceae: seeds, achenes and cypsaele
The Germination page has been reorganized with the cold stratification subject listed under Treatments for seeds germination.
Cold stratification: part 1, dormancy
Cold stratification: part 2, FAQ
Cold stratification: part 3, the practicals
Cold stratification – part 3, the practicals
Wildflowers of North AmericaCold stratification – part 1, dormancy
Cold stratification – part 2, FAQ
Let’s dive into the methods used for providing cold stratification as a seeds germination treatment, drawing from my personal experience. They often overlap with the sowing methods (also mentioned for seedlings winter protection). It makes sense because all are interrelated .
Natural cold stratification = sowing in the ground (I also call it sowing ‘in situ’)
This is the easiest method of providing cold (moist) stratification for seeds. We actually reproduce what’s happening in nature!
This type of sowing/stratification can be done until the ground freezes, so probably early November for our region would be safe to give as a limit. In case of ‘in situ’ summer sowings (Corydalis, Asarum, Hepatica…) you will ensure first a natural warm/moist and then a cold stratification.
Incorporating potting mix in the top layer of sowing will provide a better substrate for the future seedlings.
Cons: selecting a proper location is the key to this method; it should be good for the germination and further growth of the plants. In this case we usually do not aim to transplant the young seedlings.
The seedlings are prone to be easily damaged in first stages and so their mortality may be higher than in pots. Sowing seeds in larger quantity it is necessary in order to establish young plants.
Despite the cons, many wildflowers are self-seeding with success in nature, and even in the garden. Lobelia siphilitica, Asarum canadense, Corydalis, Hydrophyllum, Geum rivale, Iris versicolor, Lathyrus vernus, Primula mistassinica to mention just a few, are regularly self-seeding in my garden.
In case of hemiparasitic species it is definitely worth trying. Although I had Castilleja species germinating well in pots, a flowering plant was obtained one year only after sowing seeds directly in the garden.
Erythronium grandiflorum seedlings, in situ sowing 2021
Castilleja coccinea seedlings 2022, in situ sowing
Another nature inspired method involves Providing cold stratification in moss (outdoors).
This was shown for Gaultheria procumbens, and it involves sowing the seeds in moss, in the garden or on a piece of moss placed in a pot outdoors. The moss will provide protection and constant humidity for the small seeds and seedlings.
For those more creative, a piece of decaying stump or a boulder are good opportunities to try and establish moss and then sow suitable species.
Saxifraga cuneifolia moss sowing, 2015
Moss with Gaultheria procumbens
This method can be tried for Gaultheria, Coptis trifolia, Trientalis borealis, Maianthemum canadensis, Cornus canadensis, shade growing Saxifraga, Rhododendrons and probably many others. In nature, we always see young Hepatica americana and Gaultheria procumbens seedlings germinated in moss patches.
Cold stratification in pots or larger containers buried in the ground
In this case we sow in pots or in larger containers and then place them in the ground. We can start with the first summer sowings and continue until late fall before the ground freezes. You can use an ’empty’ area between the garden plants, or organize a special place in the garden ‘out of sight’.
Sowing in a larger container works very well for species with the same germination requirements that need to be grown in the same place for 2-3 years.
Before the winter, add a layer of dry leaves/mulch for extreme cold protection; it will also help compensate for the lack of snow (if).
Avoid low areas where a lot of water might stagnate in early spring after snow melt!
Cons: the location to keep the pots for the winter may not also be the best for the germination and growth of the seedlings; you may need to move them in the spring in another location. For this reason, many times I keep the pots buried for the winter and then move them as soon as the spring arrives. It adds as extra work to be done.
Summer sowings need constant monitoring/watering and being ‘buried’ in the ground, makes it easy to forget about them.
The seedlings are also more prone to damage in early stages.
Cold stratification in the ground, 2019
Sowings for cold/moist stratification: Diphylleia, Daphne…
Cold (moist) stratification above ground
For those who prefer the ‘cleaner’ or classic method, sow in pots/containers and provide the cold stratification above ground.
We need something to keep the pots together, hence the box method, easy to bury in the snow during the winter.
Any other container type, small or large, plastic trays, work well to keep the pots in place. Over the years I have also used plastic (nursery-type) trays with a mesh on top, but that came with the challenge of fixing the mesh. Another option is to wrap the trays in a plastic sheet, which works pretty well.
Cold stratification in a box, 2025
Cold stratification tray/plastic,2015
As previously mentioned, make your cold storage pile in a shaded area to make sure the snow remains compact and doesn’t melt during our rare sunny days. Be creative in using your garden space: underneath a patio table, a bench, beneath eavestroughs or a large evergreen tree…This way the pots will also be protected from occasional heavy rains, an important fact to keep in mind in the spring when the emerging seedlings are very fragile.
Cons: snowless winters can be problematic for protecting the pots from extreme cold.
The pots will need to be handled/moved around multiple times.
Cold frame November 2017, before adding insulation and a lid
If you want to grow a lot of plants/species, a classic frame or a simple wood framing are very valuable in our climate. They do not need to be fancy and have multiple advantages.
Place your pots inside tight together and towards late fall add some insulation along the edges: plastic sheets, bubble foil insulation…); on top better to add a thin fleece and a plastic foil. A lid is beneficial and it can be easily improvised, by making a light frame with plastic in between. This is what I used when I had my very simple, home-made frame and it worked well.
The advantage of a frame is that you can easily add bubble sheets and tarps on top of the frame to compensate for the lack of snow. You will still find the pots frozen when first opening the frame, but apparently the temperature remains acceptable. Various seeds always germinated well and all young seedlings survived.
A frame also provides a good place for keeping all your pots organized together: sowing and seedlings during the growing season (a shading cloth/anti-critters mesh to be used during the season).
Cons: a limiting factor in this case is having a proper, large enough space with good exposition for installing the frame. Avoid a south facing location because during the summer it will get too hot for small size seedlings and the pots will dry faster.
For people with mobility issues, working with a ground frame may pose problems; there is a lot of bending and crouching involved to oversee the pots.
Cold (moist) stratification provided in the fridge
In this purpose we place the seeds between moist paper towels enclosed in plastic bags, in moist media (vermiculite, perlite…) or keep whole pots in a
dedicated fridge/shelf. This method was advertised a lot as a space saving method and it looks ‘attractive’. I employed it for a while, discovering its many disadvantages in the process.
Cons: the seeds need to be inspected frequent, especially in moist towels. Even if you pay a lot of attention, you may find germinated seeds with the roots entangled in the moist towel, or even worse with etiolated shoots.
Most important, most seeds do not germinate uniformly and they would need potting up when/as it happens; damage to the rootlets when potting up is also a factor.
Poor germination results are also possible due to the constant fridge temperature, and also because of the molds sometimes forming on the seeds (when using moist towels).
Asclepias exaltata germinated seeds in/through moist towel, 2018
For these reasons I now consider this as a last method for cold stratification by those living in a cold climate. In any case, stratification in moist media is preferable than using moist towels, with the exception of very fine seeds.
I recommend the fridge stratification only if it is too late to provide the stratification outdoors, to keep seeds moist at cold for a very short period, or when a reliable method that works well has been developed, like for example: How to easily germinate Hydrastis canadensis.
***Special attention is needed for the seeds that were sown early in the season like: Hepatica, Sanguinaria, Corydalis, Jeffersonia… in fact the whole category of the ‘moist-packed’ seeds which have been discussed ad nauseam in this blog.
I found this image from November 2014! showing with my first germinated seeds of Paeonia mlokosewitchii! (roots only; the top layer of potting mix was removed to take the picture). We will see the germinating shoots in part 4.
But let’s not forget that other species also require first a lightly warm cycle followed by a cold stratification in order to germinate well: most Paeonia species, Helleborus, Actaea, Aralia, Hydrophyllum, Viburnum…
Keep in mind that in this case, by fall there are radicles/tiny rhizomes growing in the pots!!! (the lightly warm/moist cycle), even if we do not see any growth above the potting mix.
A cold stratification it is then needed for the shoots to be induced into growth. For this reason I sometimes called them ‘double stage germinators’ (easy to remember what’s happening).
The best methods for cold stratification for these species are: pots or containers buried in the ground or a cold frame.
Next in Part 4 – spring arrives: what to expect and how to handle the start of germination following all the shown methods.
July colors
Wildflowers of North AmericaLet’s have a short break from the cold stratification with flowers from July 2025, to visually counteract the white, bitterly cold outdoors. There are many plants flowering in July; usually the drought hasn’t installed yet and some of the June flowering plants extend resulting in riot of vivid colors. The seeds collection intensifies in July so there aren’t as many pictures taken like in May. But there are enough for a quick review :)
I will keep the Campanula species separate, for my annual Progress with the Campanulaceae. Phyteuma orbiculare and Physoplexis comosa also flowered in 2025 so it is going pretty well.
A special note for Campanula rotundifolia Kill. (read more here: Campanula rotundifolia complex). It started to flower in June and kept going until fall! A beautiful tall bellflower (it may need support), with ribbon-like foliage and an abundance of flowers. It can be grown from full sun to part-shaded locations.
Campanula rotundifolia (Kill.)
Campanula cochleariifolia
Campanula glomerata ssp. cervicarioides
Images arranged after location: from sun to part-shade and shade
Centaurea jankae
Platycodon grandiflorus ‘Sentimental Blue’
Zigadenus elegans ssp. glaucus
Clematis hexapetala
Echinacea
Crocosmia ‘Lucifer’
Lilium-martagon-var.-pilosiusculum-flowers
Swertia perennis
Mimulus lewisii
Lilium michiganense
Arisaema franchetianum ‘Hugo’
Asclepias exaltata
The majority of the plants shown were grown from seeds. Here’s the breaking down for the germination method:
By sowing in late fall/early winter (cold stratification): Clematis species, Delphinium fissum (seen behind the Centaurea) Swertia perennis, Asclepias exaltata and Zigadenus elegans var. glaucus.
All others: warm germinators (sow in early spring indoors if you use a light stand, otherwise sow in pots in the spring outdoors).
*Lilium species need a warm/cold stratification for complete germination.
Cold stratification – part 2, FAQ
PropagationThe post prepared for today was too long (not again! I hear) so here’s just half of it. A FAQ and few interesting facts about snow insulation.
FAQ
First, which are the plant species that require this treatment in order to germinate well? Many of them, too many to list them here. There is data available for many of the cultivated species. In our seeds offerings these species have the germination info saying: sow in late fall/early winter or, cold-moist stratification required.
How long the cold stratification has to be?
Theoretically, the length of the required cold period can vary from a couple of weeks to a few months (see the last post). In some cases, it is impossible to tell the exact length period because it depends on the type of dormancy, the temperature applied and other factors.
How cold it has to be for the cold stratification to be effective?
break and only some species (best known those from fam. Ranunculaceae, various Gentiana. probably other species as well) will actually require below 0C stratification.
It has been proven that for many species regular fridge temperature around 0-4C is OK for the dormancy
However, not all species I’ve tried in the fridge reacted to the treatment. Not only the temperature matters, but also how it is applied. In nature, the temperatures vary during the winter months, and also from day to night. We cannot ‘reproduce’ this in our home fridge and this may be the reason why not all the species react well to stratification in the fridge.
How low is too low, temperature wise?
This question has preoccupied my for quite a while because I usually sown and keep all my pots outdoors. I still do not know the answer; probably no one has gone into this level of details.
In any case, keeping in mind that these seeds are imbibed in water, and in part some physiological processes are already happening inside the seeds, we can assume that for example a range of: -10C, -15C would be too low.
Do we really need snow?
It depends on the stratification method and your location in regards with the winter minimum temperatures. Probably gardeners on the west coast do not need to worry about snow. The insulating benefits of snow are well-known, especially the fluffy kind of snow because the air that is trapped in between the flakes.
Doing a quick search AI kindly informs us that:
“Snow acts as a natural insulator, like a blanket, because its intricate crystals trap a large amount of air (90-95%), which slows heat transfer from the soil to the cold atmosphere, protecting plant roots and soil life from extreme temperature fluctuations and winds. The effectiveness depends on snow quality: fresh, fluffy snow is a great insulator, while hard, compacted snow is a poor one”.
There is also good info on various aspect like:
Depth thresholds: For significant insulation, a certain thickness is required. Measurable warming effects typically begin at 25–30 cm of snowpack. Research in 2026 suggests site-specific thresholds for ground stabilization can be as low as 4–7 cm in alpine regions.
Allium thunbergii ‘Ozawa’, Jan. 15, 2026
Albedo effect: Snow’s high reflectivity (albedo) reflects 80–90% of sunlight back into space, keeping the surface cool during the day.
Average conditions: In moderate winter climates, 10 cm of snow can maintain ground surface temperatures up to 9°C higher than bare ground.
Extreme cold: Studies show that while air temperatures might drop drastically (e.g., to -15°C), the soil under just 10 cm of snow often stays near or slightly below freezing (0°C to -5°C), creating a difference of 10°C or more.
Apparently, 10 cm of snow has an R-value of approximately R-4 (roughly R-1 per inch; comparable to the insulating value of a layer of wood or a thin layer of fiberglass insulation).
We do not always have winters so cold like this one and we are lucky to also have copious amounts of snow, just like last year. So, even if the low was -14C in our area yesterday, with the +/-30 cm of snow the temperature at ground level should have remained decent!
Apparently, even 10 cm of snow are enough to do that.
It explains why the frozen pots I always find in very early spring, either buried in the ground or in the cold frame (while I had it), were not negatively affected germination and/or seedlings growth wise.
Cold stratification, Corydalis 2019
Hepatica nobilis frozen seedlings in the cold frame
Corydalis malkensis, always the first to appear in early spring; frozen ground.
It also explains the buds apparition of the very early flowering species seemingly from the frozen ground in late February or early March, depending on the snow melt: the snowdrops, various Corydalis, Cyclamen coum, Eranthis, some peonies buds and even Helleborus.
In between the snowfalls, temporarily thaws are happening; we just had one last week. The snow melts in some degree and sheets of ice are forming. They are not good insulators and also the weight can be considerable (especially on top of pots above ground). But do not worry, over the years I noticed that there will be no damage.
Above ground container with Sarracenia purpurea two days ago and today. Drosera seeds were also sown in it in late fall – the easiest method of providing cold-moist stratification for seeds :)
Next in Part 3 – cold stratification, the practicals
Cold stratification: part 1, dormancy
PropagationHappy New Year to all again! As the saying goes: out with the old and in with the new, which also means making order in various aspects of one’s life. Website wise, besides deleting older posts with no real significance long term, more important pieces are brought together and pinned on the Germination page.
The cold (moist) stratification as a germination treatment was mentioned many times when talking about germinating various species and how to store your sowing and seedlings over the winter. However, I think one more detailed, only seeds-related post is needed.
First, briefly about seeds dormancy, mostly for those new to germinating seeds. We cannot talk about cold (moist) stratification without a few notes on dormancy.
Easy germination of various non-dormant seeds (species)
As we know, many annuals, and also various perennials, will germinate easily in proper conditions related to light, temperature and humidity. These are non-dormant seeds and we wish all were like this! Examples would fill a whole page: Silene spp., most Dianthus, Bukiniczia, Erinus,some Campanula ssp., Papaver…
On the other hand, in the same conditions, many other seeds stubbornly refuse to germinate; sometimes even after being ‘attacked’ with various so called germination treatments. This is because of the so called seeds dormancy, which we courageously attempt to ‘break’.
The seeds dormancy is commonly described as possible to be: physical, physiological in various levels, morphological and morphophysiological. Gradients between types of dormancy are possible. Many times the seeds may in fact have a combination of these: for example physical + physiological dormancy.
Without going into details (AI can help with that), the physiological dormancy is found in seeds of many species including many perennial plants, also shrubs and trees, and most often it can be broken by the use of a cold (moist) stratification period. It can be: non deep, intermediate and deep, and I found the naming self-explanatory. We can understand why some species will germinate after 1 week of cold (moist) stratification and others after 4 months (those with deep physiological dormancy). One example: Lewisia rediviva with germination after a +/- 1 month of cold stratification and Lindera bezoin with germination after 3+ months. I won’t go into details about the others, will give examples in part 2.
Something else to note is that in some cases even non-dormant seeds can become dormant due to environmental conditions; also dormant seeds can enter a second-dormancy making it even more difficult to ‘break them’.
Hepatica japonica 1 and 2 years old seedlings
Another useful bit: in a given lot of seeds, not all need to have the same type of dormancy. For example, part of the seeds can have physiological, while another part can have morphophysiological dormancy.
In translation – not all seeds will always break dormancy and germinate at the same time (examples: Glaucidium palmatum, Sanguinaria canadensis, Hepatica…).
Back to the cold (moist) stratification which is most commonly required for breaking seeds dormancy in many species, be it as a single treatment or in combination with a warm (moist) stratification (multiple cycles may be required), or in combination with other treatments.
Once upon a time, before we had a million studies about seeds dormancy and its classifications, people would sow seeds in the fall and in most cases they would germinate in some percentage in the spring. In translation – the seeds were applied a natural cold (moist) stratification during the winter.
In nature, most often seeds/fruits fall on the ground when they mature, which can happen from early summer to late fall. The ecological approach to seeds germination always takes this into account.
For example, eeeds maturing in early summer will naturally undergo a warm stratification before the fall arrives and the cold stratification starts, so it makes sense to treat those seeds in the same way when we collect and sow them. If we collect and store them under improper conditions in our home until sowing in the fall, is not the same. It should be no wonder that they react differently and maybe will not germinate in the spring. Examples: Corydalis species, also Erythroniums and other early spring flowering species.
Unfortunately, many studies on seeds germination do not follow an ecological approach. Hence, the contradictory germination results for various species. The same applies for some germination experiments done by gardeners.
Another useful info I would like to share on the subject is that after seeds dormancy was broken, different conditions may be required (for some seeds) to stimulate the germination.
I’ve read it long time ago, but it has taken time for me to really understand the concept. I noticed that some seeds (species) kept in the fridge (vermiculite or moist paper towels) will start growing roots after a while (usually toward the spring) while others not, germinating only after they were taken out the fridge.
Translation 1 – the conditions suitable for dormancy break and germination may or may not be the same, depending on the species.
Translation 2: maybe the seeds are not dormant anymore, but they won’t germinate due to unfavourable conditions.
The general idea I am trying to convey is that some of those little grains we are trying to ‘break’ out of dormancy, may have many reasons for refusing to germinate. There are still many unknowns related to the dormancy and germination, plus there are many differences between the many plant species!
Smilax tamnoides seeds, dormant – keep quiet :)
Something to meditate on until part 2 is ready:
“The dormant-seed stage in the life cycle of many plant species should be visualized as a period of time when things are happening in the seed, e.g., physiological/biochemical and morphological/anatomical changes, growth of the embryo, mobilization of food reserves and certainly activation and deactivation of genes.” (Baskin & Baskin, 2014)
Next in part 2 – we’ll get back to the practical approaches for providing the cold (moist) stratification for seeds (boxes, snow and all that…).
Happy cold-moist stratification!
PropagationHappy New Year to all! Good health, happiness, good germination, beautiful plants and gardens in 2026!
We all hope for a better year and there is nothing more hopeful than sowing seeds and waiting for their germination in the spring!
Using a plastic box is not mandatory, but it will just prevent critters scratching the pots during the snowless periods and in the early spring (a mesh fixed on top works the same).
I continue to use the term ‘cold-moist stratification’ to make the process clear and because I noticed that some people use it with a wrong meaning: “stratification” per se implies a moist medium. Keeping dry seeds in the fridge do not equal stratification.
For many seeds (species) that’s all they need to germinate properly: 2-3 months of cold (moist) stratification, preferably with variations in temperatures throughout the cycle.
Not all the species need it to be as cold as it is right now, but certainly those from the Ranunculaceae family are having a blast this winter!
*What happens if species that are actually warm germinators are sown in the fall/winter? Nothing bad, good seeds (attention to most species from fam. Asteraceae) should germinate in late spring with the raise of temperature (with the exceptions of the ‘two-stage’ germinators that first require a warm cycle followed by a cold one in order to germinate – those I keep advocating to be sown in July-August at the latest).
Short review April 2025
PropagationContinuing the review, at the beginning of April, spring was still slow to start; the Corydalis were trying to join the snowdrops without much success. Another very early
spring flowering, Physochlaina orientalis, boosted by the snowmelt, was waiting for the first pollinators.
Then, like it often happens in this area, a sudden rise of temperatures in mid April resulted in the overlapping flowering of many spring wildflowers: Corydalis, Hepatica, Helleborus, Jeffersonia dubia, Draba and Primula species, Pulsatilla….
Corydalis in their many forms and colors were showcased many times in the past, but ‘new’ for this year was a beautiful patch of Corydalis malkensis mixed with Corydalis solida purple, due to my unintentional replanting: a nice mix for our Corydalis ‘Wildheart mix’!
Their ‘relative’, Dicentra cucullaria (a cultivated form), had an exceptional good flowering this year. The white form of Jeffersonia dubia flowered for the first time, reason why I selected it for this article’s cover image.
Erythronium albidum tried to flower more than its usual, Primula glaucescens usually struggling with the not-moist enough location flowered timidly while Fritillaria collina not only flowered well, also produced seeds for the first time later in the year. From the many nice seedlings obtained from the Helleborus ‘Cherry Blossom’ it is enough to show only one :)
Physochlaina orientalis, April 2025
Pulsatilla styriaca pink, 2025
Primula mistassinica, 2025
Paeonia ex. wendelboi, 2025
Primula glaucescens
Erythronium albidum, 2025
Helleborus ‘Cherry Blossom’ seedling 2025
Corydalis ‘WM’ 2025
Hepatica americana #2, 2025
Besides plenty of snow melt, there was also rain in April, and most importantly, no late frosts coming into May! Not too many sunny days, but otherwise it was a very good start of the ‘real’ spring.
Early spring 2025 review
PropagationGalanthus elwesii and Helleborus x hybridus flowers buds in the background
Believe it or not, the year’s end is fast approaching. Looking back at the gardening season is always a reason of joy, sprinkled with little disappointments for things that didn’t go as expected. We learn and move forward!
As I said in a previous early spring review (2022), keeping pictures-records of your garden can serve many purposes besides showing off on social media.
Use them to: compare the emergence of species over the years, decide planting/transplanting locations for various species, understand which of the species are the most resilient and useful to add/propagate more for an early spring show. Every June, I gather all snowdrops capsules and carefully sow the seeds in pots or directly in the garden.
Pictures taken in the spring of 2025, show a very late start of the season, with the snow lingering late into March in parts of Ontario. In this context, all very early bloomers are always a blessing for our winter frozen souls.
Just a few images from March 2025, to better illustrate what species you should grow for early ‘signs of life’ and flowers in a similar climate. The first red shoots of various Paeonia peeking through the ground are equally valuable for me like the blooming snowdrops (Galanthus), Crocuses, winter aconites (Eranthis) and Iris reticulata (‘Katharine Hodgkin’ remains the more reliable). And what’s the very fresh looking rosette right from under the snow? A Capnoides sempervirens 😊
Paeonia mlokosewitchii, March17 2025
Eranthis, March 2025
Crocus March 17, 2025
Iris ‘Katharine Hodgkin’ 2025
Helleborus purpurascens, 2025
Capnoides sempervirens March, 2025
Perennial cold hardy Adonis species are a big bonus. The first Adonis amurensis raised from seeds has timidly bloomed this spring, keeping company to Adonis ramosa. If you manage to add them in your garden do not hesitate; it is worth waiting for them to grow up. The same goes for the tiny Cyclamen coum.
Adonis ramosa, March 27
Adonis amurensis, March 27, 2025
Cyclamen coum March 17, 2025
*This is still an excellent time to sow various species for cold/moist stratification, keep the pots in the house a bit and then bury them under the snow!
My latest sowings: Viola rostrata (in order to have more seeds we need more plants!) and Clematis alpina ssp. sibirica (wild collected seeds are precious).
Winter ready
PropagationWe are all now preparing our gardens, sowings and seedlings for the winter. I already provided a few easy solutions for storing various pots in a previous post, available on the Germination page: How to prepare your sowings and potted seedlings for the winter.
I just took few more pictures while arranging my pots with seedlings, sowings and leftover potted plants in the past days. I do not longer use a cold frame, so back to the basics: ‘planting’ the pots in the ground and/or bunching them together and later cover with a fleece/tarp. It is the best, easy method to provide the cold/moist stratification required by various species. Easy-peasy, everyone has some space in the garden between perennials and shrubs that can be used this way: sown this fall Diphylleia, Daphne mezereum and few others (species that require cold/moist stratification), pots with Hepatica and Helleborus seedlings, young, potted Thalictrum petaloideum planted for the winter.
Sowings for cold/moist stratification: Diphylleia, Daphne…
Helleborus seedlings, Hepatica
Thalictrum petaloideum
New for this fall, I am using a tall container that was filled with annuals during the season, to ‘plant’ some old and new sowings little pots. While cleaning away the petunias and others, the large empty space was begging for few pots to be buried in it. Being above the ground I don’t know how it will work but worth to give it a try .
We’ll see how it goes.
Friday’s seeds special mix
PropagationA mix bag of seeds for today. After the first snow, usually we start thinking about the stocking stuffers. These are all seeds from garden grown plants, in limited quantities, that were just added in the inventory. Maybe you will find something special for you and your gardening friends 😊in our SEED LIST.
Something for the Gentiana lovers:
Gentiana paradoxa – flowering by itself this year, so the seedlings will be true to species: Marvellous Gentiana
Plus, Swertia perennis which got a great start in late spring before the drought arrived, so
the seeds are very limited.
Gentiana paradoxa 2023
Swertia perennis
Something for those looking after really interesting rockery plants (show them to your buddies or on social media kind):
I took advantage of the bad weather and sorted out through the Bukiniczia cabulica ‘fruits’.
We discussed about Bukiniczia ‘seeds’ a few times. A very easy to germinate biennial plant, IF the seeds are good.
Physoplexis comosa – new to our SEED LIST, where you can also find other special plants.
A plant portrait and more details about this interesting rock garden species on the product page.
Bukiniczia cabulica
Physoplexis comosa 2025
For those looking to extend their fall flowering season:
Lomelosia olgae ; we already talked about it here: Friday’s seeds and a plant portrait
For the wild orchids aficionados (micropropagation ‘in vitro’):
Cypripedium acaule, the Mocassin flower.
Lomelosia olgae, October
Cypripedium acaule
Reminder:
*The priority at this time is to first sow all species that require a cold/moist stratification and organize a place for the pots outdoors.
** There are still some moist packed seeds left in the inventory, do not wait for much longer; the shipping window is getting smaller by the day!
Friday’s seed with Water arum
Plant portraits, PropagationMucilage enclosing the seeds of Calla palustris
The abundance of red fruits of the Water arum, Calla palustris, draw my attention the past weekend while looking for (and not finding) skunk cabbage fruits.
Although the last, bad experience of extracting the seeds from the Water arum berries was still vivid in my mind, I took a couple of them.
The fruits are clusters of red berries; each berry contains a few seeds, enclosed in a mucilage, which is almost impossible to get rid off. The seeds are best extracted by squeezing the end of the berry; even so, the ‘blobs’ of mucilage seem to have a long life of their own :O
The Water arum is a very hardy and reliable flowering species for any wet areas in cold climates, easily growing and forming colonies in still, shallow waters like ponds, streams edges; it can also be cultivated in large water-containers.
Like other members from the Araceae family, the ‘flowers’ are formed from a showy white spathe surrounding a cylindrical spadix bearing the greenish-white, small real flowers.
Calla palustris
Calla palustris
Calla palustris fruits
I will not repeat the experience too soon, so those with proper garden conditions do not miss this opportunity. Very important, the seeds need to be kept moist packed!
Other native species added to the inventory:
Aruncus dioicus – new
Asclepias exaltata
Latest for October
Wildflowers of North AmericaCeanothus americanus in bloom
The latest seeds collected and added/replenished in the inventory belong to the following native species:
Ceanothus americanus, New jersey tea
Desmodium nudiflorum, Naked flowered tick-trefoil
Euonymus obovatus, Running strawberry-bush and
Smilax herbacea, Herbaceous carrion flower
These are all drought resistant species and are good options for those looking to add new native species to their easy-maintenance shade garden.
The New Jersey tea in particular, is an extremely adaptable small size shrub, growing very well in poor soils, with white, fragrant, pollinator friendly flowers. You can also grab a few leaves from the garden when you run out of tea!
Add the strawberry-bush as a ground cover around it and you are done for a really simple, maintenance free plants combination!
Euonymus obovatus in late fall
The time to shine for the Running strawberry-bush is in late fall, when the green foliage becomes more visible on the woodland floor showing the conspicuous red capsules with orange seeds.
The same goes for Smilax herbacea; in good years, the foliage turns bright yellow showcasing the black fruits clusters (in the featured image).
* This fall the Hamamelis capsules were damaged by the weevil larvae (Pseudanthonomus hamamelidis).
**From other species with late fruits/seeds maturation I also hope to collect Clematis virginiana and Cephalanthus occidentalis in the near future, if possible.
Due to the extensive summer drought, most likely there won’t be any skunk cabbage fruits this fall (for those who asked).