A well done job

Never enough gentians….

In my opinion the bumble bees did a great job with Gentiana andrewsii. There are enough seeds for Canada 150 Celebration sale, and also for the seeds exchanges! :)

Bumble bee pollinating Gentiana andrewsii

A well done job

 

 

Friday Seeds & germination requirements for Viburnum species

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This fall I collected for the first time fruits of Viburnum acerifolium (maple-leaf Viburnum) and Viburnum rafinesquianum (downy arrow-wood Viburnum).

Surely, I was curious to see how the ‘seeds’ (endocarp + seed) are looking but how about their germination?
There are contradictory statements on the web; some say they first need a cold stratification, others say warm. It’s good to clarify this because I like to provide reliable info.

So, back to the books ;)

Viburnum acerifolium pits (endocarp+ seed)

Viburnum rafinesquianum pits

I like Baskin & Baskin: Seeds – Ecology, Biogeography, and Evolution of Dormancy and Germination (2014) because besides giving results from their own experiments, there are citations of hundreds of other articles on germination (of course, in some cases things are not always clear).

And, the warm stratification is the winner!
These Viburnum spp. have deep simple epicotyl morphophysiological dormancy and the germination requirements are: warm/moist stratification period (during which roots emerge), followed by a cold/moist period (after which the shoots grow).
In translation, if sown in late summer they will show up the next spring; otherwise they will germinate complete in the second spring after sowing.

On the same list are many other Viburnum species like: alnifolium, betulifolium, cassinoides, dentatum, dilatatum, lentago, opulus, parvifolium, prunifolium….Those who don’t have access to the book can read here a very short but conclusive summary regarding Viburnum acerifolium germination:
http://www.bioone.org/doi/full/10.1674/0003-0031%282005%29153%5B0232%3AEDIVAC%5D2.0.CO%3B2

Also another summary here:
https://www.researchgate.net/publication/267996605_Overview_of_seed_dormancy_in_Viburnum_Caprifoliaceae

Viburnum acerifolium

Viburnum rafinesquianum fall

Here they go!

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Trillium grandiflorum seeds are just beginning to germinate! As well is Paris quadrifolia; I noticed the first signs last Saturday (with my x40 lens).

I snapped a few pictures when I was packing some today; they are advanced enough to be noticeable even if the pictures are not great.

Trillium grandiflorum seeds starting to germinate

 

Paris quadrifolia seeds barely showing the radicle

No matter what someone else says or writes, this is happening every year.

For these species, moist stored seeds kept at warm (room temperature) will always start to germinate around this time (roots only).
From all T. grandiflorum seeds, 70-80% will germinate now (roots) and then show the first leaves in the spring after a period of cold stratification. The rest will need one more warm/cold cycle to go through the same cycle.

Paris quadrifolia seeds were also tested two years ago and at that time they all germinated by November (this year I only got few and were promised to someone).

On short, hurry up if you think about Trillium grandiflorum; right now I can select and send seeds that are just about to germinate (they are enlarged and lighter in colour – see the feature image).

 

 

Friday’s seeds – Mitchella repens

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I recently collected more partridge berries, together with Lindera benzoin and Prosartes lanuginosa (already shown, click on names).

Mitchella repens presents an interesting fruit, therefore worth a closer look. The red, fleshy, ‘two-eyed’ fruit it is formed after the fusion of the flowers ovaries – the white, fragrant flowers are disposed in pairs.  So, each pair of flowers will form one fruit. Each fruit has a spongy inside (a bit like Gaultheria) and will contain up to 8 seeds.

Mitchella repens fruits and seeds – Partridge berry, twinberry, squaw vine (Fam. Rubiaceae)

Why this perfect, evergreen ground hugging plant is seldom seen cultivated in the gardens remains a mystery to me. It can be easily propagated by cuttings as well.

Mitchella repens flowers

Mitchella repens fruits

Belated Friday’s seeds – Chamaedaphne calyculata

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Instead of writing about seeds, I was in collecting them ;) therefore a combined post about Chamaedaphne calyculata, the Leatherleaf or ground laurel.

The sole member of the genus Chamaedaphne (Ericaceae), leatherleaf is a low growing evergreen small shrub (up to 1.5 m tall). It is native to cool regions of the Northern Hemisphere, from North America, NE Europe to N. Japan, Mongolia and Siberia, where grows in all types of bogs, sedge fens, and open wetlands.
It is characteristic of mature and late stages of moss shrub communities, where it forms colonies, with rhizomes spreading in the sphagnum moss. It actually helps the installation of other species with whom is found growing, like Sarracenia, Drosera, Kalmia polifolia, Ledum groenlandica, various Vaccinium species.

The common name comes from the thick, leathery leaves, which are turning red-brown in the winter. It has a dense branching and the older stems turn gray with a fine exfoliating bark texture.
The white, bell shaped flowers on long inflorescences appear in April-July, depending on the region. Somewhat resembles Andromeda flowers, which is why Linnaeus first named it Andromeda calyculata.

Chamaedaphne calyculata, Leatherleaf

Fruits are capsules with split open and release the seeds shown below (wedged shaped, golden brown).

Chamaedaphne calyculata fruits and seeds

 It would make a great addition for small and large bog gardens. For those who have conditions and enjoy growing Ericaceous species – the seeds require cold/moist stratification and probably best to germinate them like in nature on a piece of sphagnum moss (in a pot/or ‘in situ’). I showed in a previous post how to easily germinate Gaultheria procumbens and Saxifraga cuneifolia in moss (Read here Sowing in moss).

Chamaedaphne calyculata seedlings germinated/growing on sphagnum moss (look around the Tamarack branch :)

 

 

In kindness – Spigelia

Few years ago I bought a postal card that I meant to frame but never got around to do it. In a lovely aboriginal design by Terry Starr (Tsimshian artist), hummingbirds are shown sipping nectar from red flowers.
The drawing named – In Kindness has the following note:
“Hummingbirds symbolize the act of offering peace and the generosity of sharing wealth”.

In kindness – by Terry Starr

I don’t know what the red flowers are but they remind me of Spigelia marilandica (Indian pink) and a very kind and generous friend who takes care to catch its seeds (explosive capsules) in small organza bags every year :)

Spigelia, with its red flowers sought after hummingbirds and their symbolism it is such a good fit for our Canada 150 Celebration Sale that I can’t abstain to add it to the list.

Let’s celebrate kindness for a while!

Those in need can read more about Spigelia – here.
*1 pck. limit/customer while on sale

 

Fridays Seeds – or is a fruit? part II Morina

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It’s Friday again so I continue the discussion (or better said monologue) about fruits versus seeds with Morina longifolia, commonly called Himalayan whorlflower.
If someone missed the Triosteum, see here.

What we call ‘seeds’ for Morina longifolia are also fruits (achenes). Morina is considered a warm germinator with the note that sometimes the hard teguments remain attached to the seedlings and are difficult to remove.

After being soaked in water for a few days, Morina achenes became soft and were easy to cut open. Actually, the tegument is quite thin compared to that of Triosteum, which is why I botched both seeds when trying to remove them (I couldn’t afford more fruits to test on).

But, we can have a look at the fully developed embryo I managed to extract from the second seed; notice the very large cotyledons.

Conclusion: soaking the seeds in water a few days (5-6), followed by sowing at room temperature should suffice for Morina. Before, I recommended keeping the seeds in a moist towel in the fridge for a couple of weeks, which is OK as well, but not really  necessary.
 

 

To autumn – the lusty song of fruits and flowers

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Images from out in the woods, from the garden and even along the margins of the roads, all reflecting the harmony of colours, shapes and textures that autumn brings, were flashing through my mind when cleaning fruits today.  Such an easy way to go into a lyrical mood…(I’ll keep it short ;)

“O Autumn, laden with fruit, and stained
With the blood of the grape, pass not, but sit
Beneath my shady roof…
………………………………………
Sing now the lusty song of fruits and flowers”.

                                           W. Blake – To Autumn

Euonymus obovatus and Arisaema

Arisaema flavum

Solidago canadensis -Rhus

Scabiosa caucasica

Colchicum

Cornus sericea

Cimicifuga racemosa

Polygonatum pubescens

Rosa

 

Friday’s seeds – or is it a fruit? Triosteum

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Sometimes we refer to ‘seeds’ without actually realizing that we are talking about the fruits.
This is the case of dry fruits like achenes, caryopses, samaras…, which tightly enclose the seed(s) and it would be practically impossible/or at least highly impractical to try to ‘extract’ the seeds.

So, why do we care if we sow seeds or fruits?
Knowing that we actually handle fruits, helps understanding why certain treatments are needed for the storage/germination process. Also, because in these cases we cannot see the actual seeds, we only make the assumption they are inside the fruit and will germinate in due time. Which perhaps they will, but sometimes there is no one there to do it!
To understand what I mean, you can have a look back at two notorious examples:
False assurance – Linnaea borealis
The Beauty and the Parthenocarpy – Acer triflorum

In the Caprifoliaceae family there are also species with dry fruits which we treat as seeds, like Morina and Triosteum. In both cases, they have hard teguments which play a role in retarding/obstructing the germination.

I wanted for a long time to have a closer look at Triosteum aurantiacum. I found no reliable info regarding its germination but someone who bought ‘seeds’ few years ago got back to me saying that by filing the fruits tegument, followed by GA3 and/or cold-moist stratification obtained good results.

So, I placed the fruits in warm water for a few days in an attempt to soften their teguments before I proceed to dissections. In Triosteum – and I refer here to Triosteum aurantiacum – the fruits resemble coffee beans, and only by looking at them one can imagine it will take a lot of time for them to disintegrate naturally in the soil.

Even after keeping them in water for few days I had to use a cutter to slice through them. The wall of the fruit is extremely thick and the seed is fused to it.
Here’s a plate showing the whole story in pictures:

I managed to extract one seed intact – inside, the embryo is invisible with the naked eye because of underdevelopment, pointing to the requirement of at least one period of cold/moist stratification.

This is a Triosteum aurantiacum SEED

Conclusion: these are the options to hurry up the germination for Triosteum – first, by filing the hard tegument with a metal file almost until reaching the seed, followed by cold/moist stratification 3 months.

Second, by nicking the end of the fruit (I used this myself for T. pinnatifidum), followed by cold/moist stratification for a few months. In this case, take care not to harm the seed when nicking the end and best to keep the fruits in water for a few days to somehow soften them.

A suplimentary GA3 treatment coupled with the cold/moist stratification may have some effect or not; I will try it this winter and we’ll know for sure.

About Morina next time…

 

 

 

Moist packing – what’s that?

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As I was packing seeds today, I thought that maybe some people who never bought or they buy for the first time moist packed seeds may wonder what they would get.

Sometimes I get questions as well, so although I have shown pictures in previous posts, it is good to tell again. There is no secret; seeds that are known to be hydrophilic (to make a long story short – these are seeds which need to be sown right away after collecting or if not, kept moist to preserve their viability), are cleaned from the fruits and stored in slightly moist vermiculite in Ziploc bags. And no, there is no need to spray the seeds with anything chemical.

These seeds are to be sown right away after receiving them. Right after collecting, for a short time (June) for shipping these seeds are packed in glassine envelopes and then in a plastic bag. Later, the portion of seeds is placed in smaller bags with a bit of the moist vermiculite or a small piece of moist towel.

Asarum canadense moist packed seeds for shipping

The small plastic bags are not intended for storing the seeds long term!!! (I know someone tried to do that, that’s why I mention it). If you really want to keep the seeds longer, and /or provide stratification in the fridge instead of the recommended sowing, you need to transfer them in larger bags with added moist vermiculite (slightly moist peat works as well).

These seeds are easily sown by spreading the content of the bag on top of the potting mix (no need to pick the seeds from the vermiculite). Even larger seeds are better sown this way. Cover with a thin layer of sowing mix, water and you’re done!

In case of hydrophilic seeds, if they are kept dry they will either not germinate at all, or will germinate poorly/over a longer period of time. Keeping them moist and allowed the required cycles that mimic the natural conditions will result most times, in 100% germination in the spring. It is a lot of extra work but worth doing it.

Thank you! – Merci! to all our customers

 

Overheating – Roscoea tibetica f. atropurpurea

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In this part of the world, the heat & humidity combined together can be quite unbearable. Today, we reached around 38C, with the humidex!
Even the little Roscoea tibetica f. atropurpurea couldn’t take it anymore and the first capsule bursted out in desperation. Two more to follow…

Roscoea tibetica f. atropurpurea with split capsule

I talked about this Roscoea a few times; a cute little thing and a curiosity. It shows up very late (June), produces only few flowers and it is so small that it needs a special place to be seen. Plus, the capsules split in a most unattractive way. Still, I found it charming :) Don’t you?

Roscoea tibetica f. atropurpurea – measured last year

Friday’s seed – Iris ruthenica

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Back to the Friday’s seed with an ad-hoc seeds photo session up in the mountains.
I. ruthenica has a relatively wide distribution from Eastern Europe to Asia, growing usually at subalpine and alpine elevations.
I particularly like it for the fragrant, deep blue/violet, flowers and the narrow leaves which form tufts of grassy ledges on the mountain slopes.

Iris ruthenica falls in the category of arillate irises and since the arils are drying fast, I wanted to capture them as fresh as possible, so not a very good image but it serve its purpose

Spending time on the mountain slope gazing at the blue sky and the rock walls would have been satisfying enough; weaving my hands through the grassy tufts to find the iris capsules made the moment unforgettable. The flowers pictures was taken during another trip, in a different location.