Continuing the Dobrogea I with other species which call this particular and beautiful region home.
I hope you already got a taste of it; this should make it easier when growing some of these species from seeds and also help selecting the garden location.

First a mention for two endemic species:

Campanula romanica:

Campanula romanica growing on a lichen covered rock outcrop, North Dobrogea, Romania

Centaurea jankae: a species with few populations known only to exist in Romania and Bulgaria. Good luck (and broken stems by wind/animals) made it to find a few late flowers to photograph.

Centaurea jankae

Remembering the sunny days of past summer, the flowers, the rocks and wildlife:

Stachys atherocalyx, very similar with Stachys recta that I offered seeds from Carpathian Mts.

Stachys atherocalyx

Hedysarum grandiflorum, as well just one late flowering stem was waiting to be immortalized :)

Hedysarum grandiflorum

Hedysarum grandiflorum habitat

Convolvulus cantabrica, perfect in a rockery and probably hardy in our area (Ontario) with reliable snow covered. I have a two-year old Convolvulus tragacanthodes growing in the rockery, we’ll see how it comes out from this winter.

Convolvulus cantabrica in wild habitat

Cephalaria uralensis

Cephalaria uralensis

The Dobrogean tortuga for a variation

Dobrogean tortuga, Testudo graeca

Thymus zygioides, found growing in the company of Paronychia cephalotes

Thymus zygioides

One more view from Macin Mts.

And a little lizard (Lacerta) basking in the sun

There were many other species of course; maybe more to follow when the next wave of arctic air hits us!

 

Well, the trip interruption was a bit longer that I would have wished, but here we are on the road again (see first part of the trip here). In order to beat the polar vortex that descended upon us we need sun & flowers!

As I said, the drive from Southeastern Carpathians to Dobrogea region is not very long, depending on the destination it can take 4-5 hours. The Dobrogea region (with Dobrogea Plateau) in Romania is surrounded to the north and west by the Danube River and to the east by the Danube Delta and the Black Sea. Most visitors are aiming for the Black Sea sand beaches, bird watching and/or fishing in the Danube Delta, and very few aim to enjoy the particular flora of the region.

Given its position, the climate is slightly warmer and more arid than in the rest of Romania, winters are cold but with less snow and the flora is very specific with a combination of endemic plants and various species at their geographical limits. It is a special ‘meeting’ place for species of Mediterranean, Eurasian and even Caucasian origin!

Even if in late July it is very hot and dry and many species are at the end of flowering, there is still a lot to explore. I will start with a few Caryophyllaceae to keep the continuity and then show various other species.

Dianthus nardiformis is an endemic of this region and luckily being a late flowering species it was still very showy; photographed on the dry, exposed, rocky hills of Enisala fortress and everywhere else. Striking especially when dispayed against the lichens laden rocks.
Cultivated for a long time, this Dianthus makes for a good garden plant, although not as tidy (foliage speaking) as other species, but extremely floriferous, hardy and long lived. There is an impressive specimen at Montreal Botanical Garden to testify it.

Dianthus nardiformis

On the same dry, rocky hills surrounding Enisala fortress, it was a nice surprise to find Paronychia cephalotes. I have a young plant grown from seeds of Crimean accession in the rockery, and I look forward to see it flowering. However, seeing a species ‘perform’ in the wild is always much better!
It is a mat forming species with white, papery bracts posing as ‘flowers’. It is hard to think it belongs to Caryophyllaceae at a superficial look.

Paronychia cephalotes and Thymus zygioides, Dobrogea July 2017

From the neglected genus Minuartia, I will note Minuartia adenotricha, a species that you will also found on the other side of the Black Sea, on the Crimean mountains.

Minuartia adenotricha, Dobrogea 2017

Gypsophila pallasii is a similar species with G. glomerata offered in the seeds shop (as well of Crimean origin); doing very well on the dry, rocky substrate in the company of Echinops, Artemisia, Xeranthemum, Astragalus, Allium ssp. and many others.

Gypsophila pallasii at Capul Dolosman, Dobrogea 2017

A must have stop for plant lovers in Dobrogea are the Macin Mountains located in Tulcea County. These are among the oldest mountains in Europe, formed during the Hercynian orogeny with the predominant rock – granite, but also sandstone limestone. Their specific look is given by the eroded granite formations.
Although a very, very long time ago these mountains were taller than the Carpathians; today the highest point is Ţuţuiatu/Greci Peak with an elevation of 467 m above sea level.

Macin Mts., Dobrogea, Romania

In the quite dry-looking landscape of late July, Silene compacta stood apart with its rich magenta inflorescences. It is not difficult to assume that full sun locations and super good drainage would be mandatory for its successful cultivation. 

Silene compacta in Macin Mts.

Many other interesting species grow in the particular habitats of the Dobrogean plateau.
Winter getaway – Dobrogea II

 

 

To finalize the query about the new Viola I was talking last week, pictures of seeds were taken (more detailed than usual): size, surface, colour, aril size and aspect are all useful characters for identification.

Viola cf. declinata seeds

Unfortunately, not having a term of comparison the identity is still uncertain. So, until I grow a few plants it will be offered as Viola cf. declinata.

While at it, why not see about other Viola spp. seeds? And, of course, why not have a Violaceae page in the Seeds Library? Have a look and notice the subtle differences between Viola renifolia and Viola macloskeyi seeds (these two native species are a bit hard to differentiate).

And how about a Viola category in the seeds shop? :)) On this occasion, I ‘discovered’ I forgot to add the Viola pubescens, as well as Viola jooi seeds to the inventory.

It’s so good to have everything in one place!

The Seeds Shop activity is back to normal; special thanks to those who placed orders and waited patiently!

About the Viola

Last day in the Carpathian Mts. this summer was dedicated to revisiting a special place, no seeds collecting in mind, just pure enjoyment of the scenery.
Going up a narrow rock ravine to join a narrow terrace grass trail (known as “brâna”) I snapped a few pictures, which included a Viola and then kept going. On the return, I stopped to collect Aquilegia nigricans seeds and also took on the fly some from the mentioned Viola thinking it would be nice to grow it in the garden together with Polygala vulgaris, like in nature.

Viola cf. declinata, and Polygala vulgaris – Bucegi Massif, August 2017

I didn’t know on the spot which species it was, so I should have taken proper pictures for ID. But we know about ‘should and would’….
Taking into consideration the site, flowering period and flora of the region, it is very probably to be Viola declinata.
Another option is Viola tricolor subsp. saxatilis (syn. subsp. subalpina).

Close up pictures of the seeds may have a final say on this; or not…To follow…

*Allium thunbergii ‘Ozawa’ and Gentiana cruciata seeds were added to the inventory.

Follow-up to Friday’s seeds – let’s go up the mountains and then travel to the Dobrogea region (close to the Black Sea, Romania) to see a few Caryophyllaceae.

I’ll start with Arenaria; yes, usually not overly ornamental plants but easy to please and cheerful when in flower. Arenaria ciliata shown below; another Arenaria found at high altitude is A. biflora.

Arenaria ciliata at about 1100 m alt.

Cerastium alpinum ssp. lanatum, the alpine mouse-ear, can be found at alpine level in quite a few mountain ranges, not only in the Carpathians; it grows on rock crevices, nooks formed between boulders, also on stabilized rock screes and alpine meadows. It is quite adaptable and the woolly foliage makes a nice addition to the large, white flowers; something good to try for any ‘not that advanced’ rock gardener (myself included).

Depending on the traveling time, there are always various Dianthus species to admire from the alpine level to the foothills of the mountain, like D. petraeus, D. tenuifolius, D. carthusianorum and D. glacialis, to mention just a few. Over the years I’ve presented and offered seeds of some of them, except D. glacialis.

Dianthus glacialis

Gypsophila petraea, a cushion forming alpine baby-breath, endemic for Carpathian Mts. usually grows on conglomerate boulders and crevices on the rock walls; often in localized areas. Excellent species for rock gardens and not difficult to grow based on from my previous experience; unfortunately, most of the capsules were immature and I found very few seeds.

Gypsophila petraea; in the image below with Asperula

Moehringia is an overlooked genus in my opinion. I like Moehringia muscosa for its fine texture and although I didn’t have a shaded rock crevice to offer, it did quite well this summer in my garden in a less than favourable position (‘protected’ by Primula sieboldii). The picture in the wild was taken in Barsei Massif (also part of Carpathian Mts.).

Quite a few Minuartia species grow at alpine level and lower; among them, Minuartia verna, the spring sandwort is a rewarding, easy to grow species so don’t be shy to give it a try.

Two cushion forming species of Minuartia from the Bucegi Mts. would be worth having in a rockery: Minuartia recurva and Minuartia sedoides. Especially M. sedoides looks similar to Silene acaulis, but of course that the flowers, when present, signal the different genus. Sometimes they grow interlaced with one another and it is even harder to distinguish them.
One year I must make a special effort and collect seeds of these species.

Minuartia sedoides with yellowish flowers, Silene acaulis and rosettes of Primula minima

Minuartia recurva

I previously showed and probably everyone knows Silene acaulis (first image in the gallery). I’ve also shown with other occasion the cute Silene pusilla (now growing in my garden as well :), so please browse through the gallery to get an idea of the habitats these species are growing in (click to open full size images).

Most pictures are from the Bucegi plateau (alpine level). The ‘green’ boulders’ composed of Silene acaulis, Minuartia and other cushion species, which punctuate the barren rocky areas, are in fact small plant communities.  The cushion-type plants are colonizers of these harsh habitats on rapidly draining rocky/sandy soils, and thus very important as pioneers for the installation of other alpine species.

Here’s one more good example with Minuartia sedoides and Primula minima taking good roots in the partly decomposed cushion.

Minuartia sedoides with Primula minima

I didn’t mention Sagina, Scleranthus and probably few others but it’s time to come down the mountain. From the South-Eastern Carpathian Mts. to Dobrogea region there is about a 4 hour drive; it won’t take long to get there – stay tuned…

A few images recently added to the Caryophyllaceae page on the Seeds Library all species collected this summer in the Carpathian Mts. and Dobrogea region, Romania.
In general, species from Caryophyllaceae family are easy to grow from seeds and some (for example,  think Dianthus, Silene) may even flower in the first year.

Pictures with various Caryophyllaceae showing the habitats they grow in, will follow this weekend. Stay tuned, I’ve prepared many pictures – the time for winter getaways has come!

 

Cerastium alpinum ssp. lanatum- alpine mouse-ear

Gypsophila petraea

 

Silene compacta

 

 

I’ve started to put together a list with species easy to grow from seeds as a way to provide info and encouragement to those who are at the beginning of their seeds adventures. The list will see additions as we go.

First, a short plant portrait fro Edraianthus graminifolius.

Edraianthus graminifolius, Grassy bells, belongs to the Campanula family and like its name suggests, has linear, grassy-looking leaves and large,

Edraianthus graminifolius

blue flowers, in terminal clusters in May-June. The foliage remains tight and low at about 15 cm but the flowering stems are spreading/trailing so it needs its space ;) A delightful, easy going plant, as long as grown in full sun/well drained locations. You don’t really need a rock garden for it; just keep in mind its growth habit and place it in a suitable spot.

Growing for the first time from seeds, or just want to try something easy? Try something from the list below.
‘Easy to grow from seeds’ means:
– easy to germinate & in large percentage
– seedlings that aren’t fussy to grow
– also, easy going plants in the garden, as long as provided with the required conditions.

I will avoid the so called two-stage germinators (although all you have to do is wait, for ex. Helleborus) or others that I never germinate/grown myself, also species with naturally low germination rates, those where the seedlings need special care and plants I’m not quite sure about. For sure there could be more species added to this list.

Easy species to grow from seeds
(they will germinate at room temperature or after being sown outdoors in the spring; aka. warm germinators)

Acinos alpinus
Allium thunbergii and few other Allium

Aquilegia canadensis and most other Aquilegia
Amphicarpaea


Arenaria (most species)
Arabis species

Anthemis carpatica and other Anthemis species (first, check very well the seeds!)
Arisaema triphyllum
Arisaema flavum and other Arisaema
Astragalus (with HWT)
Asphodeline lutea
Asphodeline taurica
Aurinia saxatilis
Alyssum
Baptisia australis (using the HWT)

Berkheya purpurea
Bukiniczia cabulica

Campanula, many species
Capnoides sempervirens
Caragana (HWT)

Carlina acaulis
Calycanthus floridus (after nicking the seeds)
Centaurea triumfettii, Centaurea orientalis, and other Centaurea species
Cerastium alpinum ssp. lanatum
Cercis canadensis

Cirsium spp. (some species, C. canuum)

Draba species
Delphinium grandiflorum and other species
Dianthus ssp.

Echinops crispus and other Echinops
Edraianthus graminifolius and other Edraianthus (not all species though)
Erinus alpinus
Erythranthe guttata

Gentiana cruciata, G. dahurica
other Gentiana spp. from Cruciata section
Gypsophila species

Hedysarum (using the HWT)
Hieracium lanatum

Ipomopsis rubra
Iris dichotoma
Iris domestica
Kalmia

Kernera saxatilis
Lathyrus vernus (light scarification)
Lilium formosanum var.pricei
Linaria spp.
Lychnis ssp.

Erythranthe guttata
Minuartia, some species
Mitella diphylla
Monarda
Mimulus lewisii
Oxytropis with HWT (or scarification)

Papaver species
Paronychia cephalotes
Phyteuma scheuchzeri and other Phyteuma
Platycodon

Primula japonica and other Primula spp. (some exceptions)
Pulsatilla vulgaris, other species, hybrids,
Physochlaina orientalis
Rhodiola

Most Rhododendron species
Roscoea species

Sedum ssp., Sempervivums
most Silene species
Scabiosa caucasica, other Scabiosa spp.
Scutellaria pontica and other ssp.

Stylophorum diphyllum (fresh or moist kept seeds)

Thermopsis (using HWT)
Thalictrum cf. foetidum
Thalictrum rochebruneanum and few other Thalictrum
Thymus species
Viola – some species

HWT – it means applying the hot water treatment before sowing; it can be replaced by light scarification.

Besides the seeds, a sunny image with Echinops crispus (curly Echinops) on this bitterly cold day; the red flowers of Anthyllis might warm us up.

Echinops crispus and Anthyllis vulneraria var. coccinea in the background, early June 2017

I secretly watched this globe thistle evolve from a small seedling into a beautiful clump this year; too bad I was away just when at its most glorious time. The underside of leaves and the stems are covered with a thick pubescence (shown in the featured image), a perfect adaptation for the drought; grown from seeds collected in the southern Ural Mts. (Chelyabinsk region).

Echinops crispus, July 2017

 Like in many other species from Fam. Asteraceae, some of the fruits (achenes) are empty, thus careful selecting of the seeds is necessary.

Echinops crispus fruits and seeds

Echinops name from the Greek words ‘ekhinos’ = hedgehog, sea urchin and
‘ops’ = face, head (referring to the appearance of the inflorescences)

I’m late with preparing the seedex donations; just sent a small one to the SCRG and since the local mail doesn’t take long, I’ll pack a few more for ORG & HP Society this weekend.

It’s not easy to decide what to give; I always start to think what would be more attractive from what I have surplus, then some need more cleaning than others, and so on…Speaking of cleaning, nothing is more disappointing than receiving the wonderful little packets only to find inside ‘not so good seeds’, or just parts of them (it happened a few times).

Special attention is needed to all species in Fam. Asteraceae because by as a rule some of the seeds are always empty and it is not always easy to see which ones with the naked eye; a strong hand lens is necessary.

For example, let’s look at Anthemis carpatica ssp. pyrethriformis I just prepared for ORGS & HP. This subspecies, endemic to Eastern Carpathian Mts., while bearing the same large flowers is smaller in size than the regular A. carpatica.
It is super easy to grow from seeds (warm germinator) and is a good plant for any rockery (attention when planting the grown seedlings in the garden – they need very good drainage). The seedlings shown are from a previous seed collection I did few years ago.

Anthemis carpatica ssp. pyrethriformis seeds

and the non viable ‘seeds’

Anthemis carpatica ssp. pyrethriformis – non-viable seeds; when shown already separated, it seems easy to distinguish them but actually I had to use a x 40 hand lens to do it!

Another Asteraceae donated to both SRGC and ORG & HP – Jurinea mollis seeds are shown in the featured image.

Two years in a row, instead of seeds I found in the seedex packets just the pappus of a dwarf Jurinea species (from AGS Seedex).
This is how Jurinea seeds look like, with variations depending on the species and the moment when the seeds were collected (the pappus can be reduced in size or it may have fallen out).

Jurinea mollis seeds

This fall I collected for the first time fruits of Viburnum acerifolium (maple-leaf Viburnum) and Viburnum rafinesquianum (downy arrow-wood Viburnum).

Surely, I was curious to see how the ‘seeds’ (endocarp + seed) are looking but how about their germination?
There are contradictory statements on the web; some say they first need a cold stratification, others say warm. It’s good to clarify this because I like to provide reliable info.

So, back to the books ;)

Viburnum acerifolium pits (endocarp+ seed)

Viburnum rafinesquianum pits

I like Baskin & Baskin: Seeds – Ecology, Biogeography, and Evolution of Dormancy and Germination (2014) because besides giving results from their own experiments, there are citations of hundreds of other articles on germination (of course, in some cases things are not always clear).

And, the warm stratification is the winner!
These Viburnum spp. have deep simple epicotyl morphophysiological dormancy and the germination requirements are: warm/moist stratification period (during which roots emerge), followed by a cold/moist period (after which the shoots grow).
In translation, if sown in late summer they will show up the next spring; otherwise they will germinate complete in the second spring after sowing.

On the same list are many other Viburnum species like: alnifolium, betulifolium, cassinoides, dentatum, dilatatum, lentago, opulus, parvifolium, prunifolium….Those who don’t have access to the book can read here a very short but conclusive summary regarding Viburnum acerifolium germination:
http://www.bioone.org/doi/full/10.1674/0003-0031%282005%29153%5B0232%3AEDIVAC%5D2.0.CO%3B2

Also another summary here:
https://www.researchgate.net/publication/267996605_Overview_of_seed_dormancy_in_Viburnum_Caprifoliaceae

Trillium grandiflorum seeds are just beginning to germinate! As well is Paris quadrifolia; I noticed the first signs last Saturday (with my x40 lens).

I snapped a few pictures when I was packing some today; they are advanced enough to be noticeable even if the pictures are not great.

Trillium grandiflorum seeds starting to germinate

 

Paris quadrifolia seeds barely showing the radicle

No matter what someone else says or writes, this is happening every year.

For these species, moist stored seeds kept at warm (room temperature) will always start to germinate around this time (roots only).
From all T. grandiflorum seeds, 70-80% will germinate now (roots) and then show the first leaves in the spring after a period of cold stratification. The rest will need one more warm/cold cycle to go through the same cycle.

Paris quadrifolia seeds were also tested two years ago and at that time they all germinated by November (this year I only got few and were promised to someone).

On short, hurry up if you think about Trillium grandiflorum; right now I can select and send seeds that are just about to germinate (they are enlarged and lighter in colour – see the feature image).

 

 

It’s Friday again so I continue the discussion (or better said monologue) about fruits versus seeds with Morina longifolia, commonly called Himalayan whorlflower.
If someone missed the Triosteum, see here.

What we call ‘seeds’ for Morina longifolia are also fruits (achenes). Morina is considered a warm germinator with the note that sometimes the hard teguments remain attached to the seedlings and are difficult to remove.

After being soaked in water for a few days, Morina achenes became soft and were easy to cut open. Actually, the tegument is quite thin compared to that of Triosteum, which is why I botched both seeds when trying to remove them (I couldn’t afford more fruits to test on).

But, we can have a look at the fully developed embryo I managed to extract from the second seed; notice the very large cotyledons.

Conclusion: soaking the seeds in water a few days (5-6), followed by sowing at room temperature should suffice for Morina. Before, I recommended keeping the seeds in a moist towel in the fridge for a couple of weeks, which is OK as well, but not really  necessary.